1) chr	: chromosome
2–3) ref_start, ref_end	: GRCh38 locus positions (0-based, half-open)
4) ref_allele	: reference allele
5–6) hap1_allele/hap2_allele	: haplotype 1/2 allele
7) experiment_accession	: ENCODE experiment ID
8) donor	: EN-TEx individual
9) tissue	: tissue
10) assay	: assay
11–14) cA/cC/cG/cT	: number of reads with A/C/G/T
15) ref_allele_ratio	: number of reads with reference / total number of reads
16) p_betabinom	: p values calculated from the beta-binomial test
17) imbalance significance imbalanced site.	: ‘1’ passes the FDR10% threshold, ‘0’ not a significantly