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. 2023 Apr 22;14:2333. doi: 10.1038/s41467-023-37960-5

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© The Author(s) 2023

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 7 — a, The orientation of homotypic TFBSs influences expression levels (left panel) and the magnitude of the effect is proportional to the number of copies (middle panel). The proportion of TFBS copies in each orientation is associated with expression levels (right panel). b, The order of heterotypic TFBSs and their orientation influence expression levels. Multiple consecutive copies of heterotypic TFBSs show larger differences.