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. 2023 May 29;13(11):1802. doi: 10.3390/ani13111802

Table 3.

List of behaviours, definitions and the period in which behaviours were collected.

Behaviour Definition Period
Immatures with group members other than their mother
Grooming Manipulation of another group member’s fur by the immature or manipulation of the immature’s fur by another group member. August 2021 to June 2022
Playing Exaggerated spontaneous behaviour by the immature and at least another individual, including object play, acrobatic play, sparring and wrestling.
Cofeeding Feeding involving the immature and another group member within a 1 m proximity in the same tree.
Immatures with their mothers
Carrying Maternal displacement, while the immature clings to the mother’s back. February to June 2022
Nursing The immature holding the mother’s nipple in their mouth.
Playing Exaggerated spontaneous behaviour, including object play, acrobatic play, sparring and wrestling.
Touching The mother placing the palm of one or both hands on the offspring’s body.
Grooming Manipulation of the immature’s fur by the mother.
Proximity The immature and the mother being within a 1 m proximity of each other.