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. 2023 May 31;13:1206111. doi: 10.3389/fcimb.2023.1206111

Table 2.

oHSVs targeting GSCs. Genome structure, activity, and GSC models.

Name Genetic alterations GSC Ref In vivo model Main results Ref
RAMBO -γ34.5Δ
-Armed with Vasculostatin (Vstat120), under IE4/5pro
(Wakimoto et al., 2012) MGG23 implanted in immuno-deficient mice -RAMBO extended median survival.
-Vstat120 synergized with bevacuzumab and reduced migration and invasion.
(Tomita et al., 2019)
VAE -γ34.5Δ, ICP6-,
-Armed with endostatin-angiostatin fusion
(Zhang et al., 2014) GBM-SC implanted in nude micee -VAE extended median survival over parental virus or recombinant endtostatin.
-Decreased microvessel density.
(Zhang et al., 2014)
MG18L ICP6-, LacZ+, Us3Δ (Wakimoto et al., 2009; Wakimoto et al., 2012) BT74, MGG4 or MGG31 implanted in athymic mice -PI3K/Akt inhibitor + MG18L kills GSCs not astrocytes.
-PI3K/Akt inhibitor + MG18L prolongs survival to 50% cures.
-Combination of PARPi + MG18L kills PARPi-sensitive and -resistant GSCs in vitro and extends median survival.
-Combination of TGF-β inhibitors + MG18L increase recurrent GSC killing.
-TGF-β inhibitors + MG18L result in 60% cured mice.
(Kanai et al., 2011; Esaki et al., 2017; Ning et al., 2017)
Δ68H-6 γ34.5 BBD deleted, ICP6, LacZ+ (Wakimoto et al., 2009) MGG4 implanted in athymic mice -Δ68H-6 increased survival.
-Δ68H-6 is safe.
(Kanai et al., 2012b)
NG34 -γ34.5Δ, ICP6.
-hGADD34 driven by the nestin enhancer-hsp68pro
(Nakashima et al., 2018) G35 implanted in athymic mice -more cytotoxic in hGSCs than rQNestin34.5, but less neurotoxic.
-extended survival of hGSC-bearing mice.
(Nakashima et al., 2018)
G47Δ-mIL12 -γ34.5Δ, ICP6, ICP47Δ, LacZ+.
-Armed with mIL12 under HCMVpro.
(Cheema et al., 2013) 005 implanted in C57/BL6 mice -antiangiogenic activity in vitro and in vivo.
-Extends survival of mice with 005 tumors, dependent on T cells.
(Cheema et al., 2013)
G47Δ-mAngio -γ34.5Δ, ICP6, LacZ+, ICP47/Us11proΔ,
-Armed with murine angiostatin (mAngio)
(Wakimoto et al., 2009) MGG4 implanted in athymic mice -G47Δ-mAngio prolonged mouse survival.
-G47Δ-mAngio combined with G47Δ-mIL12 increased survival, virus spread, and decreased macrophages.
(Zhang et al., 2013)
OV-Cmab-CCL5 -γ34.5Δ, ICP6−,GFP+
-Armed with Cetuximab-CCL5 fusion protein under IE4/5pro.
(Uchida et al., 2013) GBM30-FFL implanted in NSG mice -Multi-mechanistic efficacy in immuno-deficient and -competent
-Increased migration of NK, macrophages, CD4+ and CD8+T cells.
-OV-Cmab-CCL5 improves median survival 2.4-fold.
(Tian et al., 2022)
OV-αCD47-G1 -γ34.5Δ, ICP6, GFP+
-Armed with αCD47-IgG1 under IE4/5pro.
(Xu et al., 2021) GBM43 implanted in athymic mice OV-αCD47-G1 treatment in vivo releases αCD47 into the TME and prolongs survival. (Xu et al., 2021)
OV-IL15C -γ34.5Δ, ICP6,
GFP+
-Armed with hIL15-IL15Rα sushi domain under IE4/5pro.
(Uchida et al., 2013) GBM30 implanted into NSG mice -hGSC killing by NK cells treated with conditioned media.
-extended survival with hCD8+ T cells and with EGFR-CAR NK.
(Ma et al., 2021)
oHSV-TRAIL -γ34.5Δ, ICP6, LacZ+, ICP47/Us11proΔ
-Armed with TRAIL under IE4/5pro.
(Wakimoto et al., 2012; Esaki et al., 2017) MGG23 and MGG31 implanted in athymic mice oHSV-TRAIL increased survival in mice with TMZ-resistant primary and recurrent GSCs (Jahan et al., 2017)
OV-CDH1 -γ34.5Δ, ICP6, GFP+
-Armed with e-cadherin (CDH1) under IE4/5pro.
(Uchida et al., 2013) GBM30 implanted in athymic mice OV-CDH1 mediated enhanced viral spread and increased NK infiltration. (Xu et al., 2018)
KNE -gB:NT, scFv EGFR-retargeted gD Δ224-38. (Uchida et al., 2013) GBM30 implanted in athymic mice -Antitumor efficacy in vivo, >50% long-term survivors.
-safe in the brain of nude mice.
(Uchida et al., 2013)
KGE4:T124 -γ34.5Δ, ICP6
-4x miR-124 target sequence in 3’UTR of ICP4
-EGFR-retargeted gD.
(Uchida et al., 2013) GBM30 implanted in BALB/c athymic mice miR-124T sites in ICP4 gene did not affect antitumor efficacy (Mazzacurati et al., 2015)
R-LM113 -scFv HER2-retarged gD
-wild-type backbone
(Calzolari et al., 2008) -mGBM-HER2 implanted in C57/BL6 mice.
-BALB/c-HGG-HER2 in BALB/c mice.
Antitumor efficacy in immunocompetent mouse models. (Gambini et al., 2012; Reisoli et al., 2012)
R-613 -scFv EGFRvIII-retargeted gD Δ6-38
-wild-type backbone.
(Mazzoleni et al., 2010) L0306 implanted in NOD/SCID mice -R-613 infects EGFRvIII+ GSCs and spreads in vitro and in vivo.
-Early, but not late treatments increase mice survival.
(Appolloni et al., 2021)
R-115 -scFv HER2-retarged gD.
-wild-type backbone.
-Armed with mIL12 under HCMVpro.
(Calzolari et al., 2008) -mHGGpdgf-hHER2 implanted in C57/BL6 mice. -Significant improvement of overall median survival, but not different than R-LM113.
-30% of mice cured.
(Alessandrini et al., 2019)
oHSV/Nb-gD -γ34.5Δ, ICP6, GFP+, ICP47Δ.
-Nanobody-hCXCR4 retargeted gD.
(Sanchez Gil et al., 2022) T033 implanted in athymic mice -Infected CXCR4-expressing GSCs.
-Treatment of mice with T033 tumors did not extend survival.
(Sanchez Gil et al., 2022)

BBD, Beclin-1 binding domain; CM, conditioned media; MMP9, Matrix Metalloproteinase 9; N/A, not applicable; NSG, NOD/SCID/IL2rg; PARPi, Poly-ADP-ribose polymerase inhibitor; ULBP3, UL16 binding protein 3; VEGF, vascular endothelial growth factor.