Table 1: Human MUC Gene Family.

Ordered by chromosomal location

Gene Symbol	Cytogenetic Band	Expressing Tissue(s) at the Ocular Surface; Protein Presence in Tears
Secreted Mucins (gel-forming)
MUC2	11p15.5	conjunctiva (RT-PCR only) (1,2); tears (low level) (3)
MUC5AC	11p15.5	Conjunctiva (1,2,4,5); goblet cells (3,5); lacrimal duct goblet cells (6); tears (3)
MUC5B	11p15.5	lacrimal gland (RT-PCR only) (7); not in tears (3)
MUC6	11p15.5
Secreted Mucins (soluble)
OVGP1 (MUC9)	1p13.2
MUC7	4q13.3	conjunctival epithelia (2,4,7); lacrimal gland (7); not in tears (3)
MUC19	12q12
MUC8	12q24.33
Membrane-Associated Mucins
MUC1	1q22	corneal & conjunctival epithelia (2,4,8); lacrimal gland (6,7); tears (3)
MUC13	3q21.2	conjunctival epithelium (RT-PCR only) (2,4)
MUC4	3q29	conjunctiva, much less in corneal epithelium (2,4,5,9); lacrimal gland (7); tears (3)
MUC20	3q29	corneal & conjunctival epithelia (4); not in tears (4)
EMCN (MUC14)	4q24
MUC21	6p21.33	corneal epithelium and lacrimal gland (this paper)
MUC22	6p21.33	corneal epithelium and lacrimal gland (this paper)
MUC3B	7q22
MUC3A	7q22.1
MUC12	7q22.1
MUC17	7q22.1	conjunctival epithelium (RT-PCR only) (2)
MUC15	11p14.3	conjunctival epithelium (RT-PCR only) (2,4)
MUC16	19p13.2	corneal & conjunctival epithelia (2,4,10); mucin granules of conjunctival goblet cells (11); lacrimal gland (12); tears (3)

Citations

^1.(McKenzie et al., 2000)

^2.(Corrales et al., 2009)

^3.(Spurr-Michaud et al., 2007)

^4.(Woodward and Argueso, 2014)

^5.(Inatomi et al., 1996)

^6.(Paulsen et al., 2004)

^7.(Jumblatt et al., 2003)

^8.(Inatomi et al., 1995)

^9.(Pflugfelder et al., 2000)

¹⁰. (Argueso et al., 2003)

^11.(Gipson et al., 2016)

^12.(Jager et al., 2007)