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. 2023 May 10;136(14):1642–1652. doi: 10.1097/CM9.0000000000002678

Table 1.

Summaries of studies applying scRNA-seq to osteoimmunology research in RA.

Study subjects Source of cells Number of subjects Cell analyzed Findings Method used Date published Reference
CIA mice and RA patients Synovial tissue RA patients (n = 23) Murine CX3CR1hiLy6CintF4/8+I-A+/I-E+ macrophages (AtoMs)and human CX3CR1+HLA-DRhiCD11c+CD80-CD86+ cells AtoMs are the osteoclast precursor-containing population in the pannus tissue, and FoxM1 constitutes a potential target for RA treatment. scRNA-Seq November 18, 2019 [56]

RA patients

OA patients

Healthy donors

Synovial tissue and blood

RA patients (n = 18)

OA patients (n = 14)

HBEGF+ inflammatory macrophages These HBEGF+ macrophages may enhance the tissue destructive capacity of fibroblasts. scRNA-Seq May 8, 2019 [58]
Early/active RA, treatment-refractory/active RA and RA in sustained remission, healthy donors Synovial tissue RA patients: naïve to treatment (n = 45), treatment-resistant (n = 31), in sustained clinical and ultrasound remission (n = 36) MerTKpos TREM2high and MerTKposLYVE1pos macrophage The relative proportions of these cells in RA may be a marker to predict the state of persistent remission vs. disease flare.

scRNA-seq, multiparameter

Flow cytometry, immunofluorescent staining

June 29, 2020 [59]
Chinese subjects underwent hip replacement surgery Bone marrow n = 2 LEPRhiCD45lowBM-MSCs These cells had the capacity to differentiate into osteocytes, chondrocytes, adipocytes, and terminal-stage quiescent cells. Some new markers for purification of human BM-MSC were identified. scRNA-seq October 11, 2021 [63]
Mice Bone marrow NA Mesenchymal progenitors at different stages We find a unique cell type that expresses adipocyte markers but contains no lipid droplets. They play critical roles in maintaining marrow vasculature and suppressing bone formation. scRNA-seq April 14, 2020 [64]
RA and OA patients Synovial tissue n = 51 T cells, B cells, monocytes, and fibroblasts We identified 18 unique cell populations and defined distinct subsets of CD8+ T cells characterized by GZMK+, GZMB+, and GNLY+ phenotypes. scRNA-seq, mass cytometry, bulk RNA-seq, and flow cytometry May 6, 2019 [27]
RA patients PBMC and Synovial tissue

ACPA+RAn = 10

ACPA-RA

n = 10

CD45+hematopoietic cells Up-regulation of CCL13, CCL18 and MMP3 in myeloid cell subsets of ACPA-RA; a lack of HLA-DRB5 expression and lower expression of cytotoxic and exhaustion related genes in the synovial tissues of ACPA-RA; DRB1/DRB5 conveys an increased risk of developing active disease in ACPA+RA scRNA-seq, Immunohistoche mical staining August 17, 2021 [87]
RA patients and healthy indifiduals Peripheral blood n = 4 RA-CCPPOS B cells The expression of IL15R is enriched in citrulline-specific B cells within RA patients were capable of producing AREG RNA-sequencing, aptamer-based SOMAscan assay, flow cytometry February 23, 2019 [88]
RA patients Synovial tissue n = 5 Fibroblast 13 transcriptomically distinct clusters were revealed. Previously uncharacterized fibroblast subpopulations were identified and their spatial location within the synovium was discerned. A 3D-printed, low-cost droplet microfluidic control instrument February 23, 2018 [91]
RA and OA patients Synovial tissue NA Synovium fibroblast Seven fibroblast subsets with distinct surface protein phenotypes were identified. Bulk transcriptomics of targeted subpopulations and single-cell transcriptomics February 23, 2018 [92]
Mouse models of resolving and persistent arthritis Synovial tissue NA FAPα+ synovial fibroblast The location and function of two FAPα+ synovial fibroblasts. Single-cell transcriptional technologies May 29, 2019 [93]
Mice Synovial tissue NA Fibroblast Expression of the synovial lining markers gradually changes along the continuum of synovial fibroblast states. Endothelium-derived Notch signaling is a potential pathway for driving the differentiation of THY1 expressing fibroblasts. Trajectory transcriptional analyses, single-cell RNA-seq datasets July 2020 [94]
RA patients and mice Synovial tissue NA Fibroblast NOTCH3 signaling drives both transcriptional and spatial gradients in fibroblasts and plays an important role in inflammation. scRNA-seq and synovial tissue organoids April 22, 2020 [95]

BM-MSC: Bone marrow derived mesenchymal stem cell; CIA: Collagen-induced arthritis; NA: Not applicable; OA: Osteoarthritis; PBMC: Peripheral blood mononuclear cells;RA: Rheumatoid arthritis; scRNA-Seq: single cell RNA sequencing; TH1: T helper 1.