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. 2023 Jul 5;13:1186659. doi: 10.3389/fonc.2023.1186659

Table 2.

Current trial phase and validated targets information on copper chelators and copper ionophore for HCC.

Type Copper regulator Abbreviation Molecular formula Trial Phase Related targets Refs
chelator Tetrathiomolybdate TM MoS4 Animal model NF-kB (86, 116)
Cell and animal models MEK1/2 (48, 117)
Cell model IL-1 (118, 119)
Cell model FGF1 (118, 119)
Animal model, Phase II trial VEGF (120122)
Cell and animal models, Phase II trial LOX (116, 123)
Animal model CCO (88)
Phase II trial IL-6 (121, 122)
Phase II trial IL-8 (121, 122)
Phase II trial bFGF (121)
chelator Trientine TETA C6H18N4 Cell model IL-8 (124)
Animal model CD31 (102)
Cell model P38 MAPK (97)
chelator D-penicillamine DPA C5H11NO2S Animal model LOX (125)
chelator Choline tetrathiomolybdate ATN-224 [(CH3)3NCH2CH2OH]2[MoS4] Cell model SOD1 (126)
chelator Tetraethylenepentamine pentahydrochloride TEPA C8H23N5·5HCl Cell model PD-L1 (56)
Cell model HIF-1 (87)
ionophore Elesclomol ES C19H20N4O2S2 Cell model GSH (106)
Cell and animal models FDX1 (65, 69)
ionophore Disulfiram DSF C10H20N2S4 Cell model ALDH (127)
Cell model ubiquitin protein pathway (112)
Animal model NF-kB (115, 128)
Cell model P38 MAPK (113)
ionophore Clioquinol CQ C9H5ClINO Cell model Proteasome (129)