TABLE 4.
Motifs in the central domain of NtrCa
Binding | Hydrolysis | Hydrolysis | Hydrolysis | Binding |
---|---|---|---|---|
G173 | S207/E208 | D239 | R294 | G355/R358 |
Walker A, binds β phosphate | Switch I effector | Walker B/switch II, coordinates Mg2+ | Catalytic residue? | Binds nucleotide base |
GESGTG173KE | ES207ELFGHEKGAFTGA | GGTLFLD239EIG | FR294EDLfhRLNV | WPG355NVR358qLEN |
For the central domains of NtrC proteins of S. typhimurium, E. coli, K. pneumoniae, Proteus vulgaris, Rhizobium meliloti, Bradyrhizobium parasponiae, and Agrobacterium tumefaciens (15, 43). The motifs listed are also referred to as conserved regions 1, 3, 4, 6, and 7, respectively, among activators of ς54-holoenzyme (36, 43). Conserved region 6 begins with residue 275 of NtrC and extends through residue 312. For cases in which fewer than four of the five NtrC sequences are identical, the residue is that for the protein from S. typhimurium and is in lowercase. The numbers indicate positions at which amino acid substitutions in the NtrC protein of S. typhimurium were studied in this work.