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. 2023 Jul 6;17:1223777. doi: 10.3389/fnins.2023.1223777

Table 2.

Studies of the applications of CRISPR/Cas9-mediated genome editing in ALS.

Target Point mutation Organism/cell line Delivery Results References
SOD1 A272C Human ALS patient-derived iPSCs Electroporation Discovered early biomarkers and pathways dysregulated in ALS (Wang et al., 2017)
L144FVX Human ALS patient-derived iPSCs Adenoassociated Virus Vector Over half of the iPSCs targeted the Src/c-Abl signaling pathway, suggesting that Src/c-Abl may be a potentially useful target for developing new drugs to treat ALS (Imamura et al., 2017)
E100G Human ALS patient-derived iPSCs Nucleofection Identified activated ERK and JNK signaling are critical drivers of neurodegeneration in mutant SOD1 motor neurons (Bhinge et al., 2017)
G93A SOD1-G93A transgenic mouse Adenoassociated Virus Vector Disruption of mutant SOD1 enhances the survival of spinal cord motor neurons and improves motor function and life span (Gaj et al., 2017)
G93A SOD1-G93A transgenic mouse Use an AAV vector and inject it into the lumbar subarachnoid space Deleting mutant SOD1 via CRISPR/Cas9 prolongs survival in an ALS mouse model (Duan et al., 2020)
G93A SOD1-G93A transgenic mouse Intrathecal injection The mouse treated by split-intein CRISPR base editor had a reduced rate of muscle atrophy, improved neuromuscular function, and up to 40% fewer SOD1 immunoreactive inclusions (Lim et al., 2020)
G93A Human SOD1-G93A missense mutation iPSCs Sendai virus-based vector The mutant motor neurons accumulated misfolded and aggregated forms of SOD1 in cell bodies, causing axonopathy and aberrant neurotransmission (Kim et al., 2020)
G93A SOD1-G93A transgenic mouse Intracerebroventricular injection Rescues motor function deficits and extends survival in a SOD1-ALS mouse model (Chen et al., 2022)
C9ORF72 Deleted GGGGCC Human ALS patient-derived iPSCs Cloned into px300 plasmid and introduced to the iPSCs by nucleofection Provides a proof-of-principle for using CRISPR-Cas9-mediated excision of the pathogenic C9orf72 repeat expansion as a therapeutic strategy in ALS (Lopez-Gonzalez et al., 2016)
Deleted GGGGCC Human ALS patient-derived iPSCs Nucleofection The mutations lead to increased Ca2+-permeable and enhance selective MNs’ vulnerability to excitotoxicity (Selvaraj et al., 2018)
Deleted GGGGCC C9ORF72 patient-derived iPSCs Sendai virus-based vector Performed an extensive phenotypic characterization of ALS-iPSCs-derived MNs (Dafinca et al., 2016)
Deleted GGGGCC C9ORF72 patient-derived iPSCs Nucleofection Proved partial inhibition of an overactivated DNA repair pathway suppresses a cell death pathway is the pathology of ALS (Lopez-Gonzalez et al., 2019)
Deleted GGGGCC C9ORF72 patient-derived iPSCs Adenoassociated Virus Vector 9 The CRISPR/Cas9-mediated genome correction reduced RNA foci, poly-dipeptides, and haploinsufficiency, major hallmarks of ALS (Meijboom et al., 2022)
FUS R521H FUS patient-derived iPSCs Nucleofection Mutations in FUS result in apparent defects in MNs derived from FUS-ALS patients (Guo et al., 2017)
G1566A FUS patient-derived iPSCs Electroporation Discovered early biomarkers and pathways dysregulated in ALS (Wang et al., 2017)
H517Q Human ALS patient-derived iPSCs Nucleofection Identified activated ERK and JNK signaling as critical drivers of neurodegeneration in mutant FUS MNs (Bhinge et al., 2017)
R521H, P525L Human ALS patient-derived iPSCs Nucleofection Uncovered a pathway of defective DNA ligation in FUS-linked ALS (Wang et al., 2018)
R521H, P525L Human ALS patient-derived iPSCs Nucleofection Metabolic dysfunction is not the underlying cause of the ALS-related phenotypes previously observed in these MNs (Vandoorne et al., 2019)
R524S, P525L C. legan knockin models Nucleofection Autophagy dysfunction likely contributes to protein homeostasis and neuromuscular defects in ALS FUS knockin animals (Baskoylu et al., 2022)
TARDBP M337V TARDBP-M337V patient-derived iPSCs Nucleofection The abnormal function of BDNF may explain the aberrant TDP-43 activity (Tann et al., 2019)
M337V TARDBP-M337V patient-derived iPSCs Sendai virus-based vector The MNs with TARDBP mutations impaired mitochondrial Ca2+ uptake contributes to glutamate excitotoxicity (Dafinca et al., 2020)

ALS, amyotrophic lateral sclerosis; MNs, motor neurons; iPSCs, induced pluripotent stem cells.