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. 2023 Jul 14;17(9):1358–1368. doi: 10.1038/s41396-023-01467-0

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© The Author(s) 2023

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 4 — Yellow represents genomic features encoded by all AOA. Blue indicates metabolisms not shared by all AOA. AMO ammonia monooxygenase, QRED quinone reductase, HURM hydroxylamine ubiquinone redox module, PQQ pyrroloquinoline quinone, DH dehydrogenase, ADH alcohol dehydrogenase, Amt ammonium transporter, UT urea transporter, SSS solute/sodium symporter, Ure urease, N/C hyd nitrile/cyanide hydratases, Cyn cyanase, TCA tricarboxylic acid cycle, HP/HB hydroxypropionate/hydroxybutyrate cycle, CA carbonic anhydrase, Hyd hydrogenase, GDH glutamate dehydrogenase, GS-GOGAT glutamine synthetase-glutamate synthase, ROS reactive oxygen species.