Parameter	Description
$r_{on}$	Activation rate of an allele
$r_{off}$	Inactivation rate of an allele
$r_{prod}$	Production rate of mRNA from an active allele
$r_{deg}$	Degradation rate of mRNA
$r_{add}^{NITC}$	Additional activation of an allele (A or A’) upon nonsense-induced transcriptional compensation caused by product $A_{nonsense}$
$r_{add}^{A,B},r_{add}^{A^{\text{'}},B}$	Activation of B by A or A’ to one of two respective B active states specified in the model
$d$	Factor by which the mRNA production rate of B is lower in the A’-directed
	B active state than in A-directed B active state
$n$	Hill coefficient
$k$	Dissociation constant of the Hill function