Table 2.

Dimensionless Model Parameters

Symbol	Definition	Description	Estimated Valuea
$\mathrm{F}$	${\mathrm{n}}_{\mathrm{m}}{\mathrm{F}}_{\mathrm{m}}/{\mathrm{n}}_{\mathrm{c}}{\mathrm{F}}_{\mathrm{b}}$	myosin activity parameter	$1.38$
$\tau$	${\mathrm{k}}_{\mathrm{o}\mathrm{n}}/{\mathrm{k}}_{\mathrm{o}\mathrm{f}\mathrm{f}}$	clutch kinetic parameter	$1$
$\omega$	${\mathrm{v}}_{\mathrm{u}}{\kappa }_{\mathrm{c}}/{\mathrm{F}}_{\mathrm{b}}{\mathrm{k}}_{\mathrm{o}\mathrm{f}\mathrm{f}}$	dimensionless myosin unloaded velocity	$140$
$\mathrm{K}$	${\kappa }_{\mathrm{s}}/{\mathrm{n}}_{\mathrm{c}}{\kappa }_{\mathrm{c}}$	dimensionless substrate stiffness	$\sim \left[{10}^{-5}-{10}^{-1}\right]$
$\theta$	$\theta$	clutch reinforcement strength	adjusted
$\Delta$	${\mathrm{F}}_{\mathrm{t}\mathrm{h}}/{\mathrm{F}}_{\mathrm{b}}$	dimensionless clutch reinforcement threshold force	$2.9$
${\mathrm{K}}_{\mathrm{off}}$	${\mathrm{k}}_{\mathrm{o}\mathrm{f}\mathrm{f}}^{\mathrm{c}\mathrm{a}\mathrm{t}\mathrm{c}\mathrm{h}}/{\mathrm{k}}_{\mathrm{o}\mathrm{f}\mathrm{f}}$	dimensionless catch bond dissociation kinetic rate constant	adjusted
${\chi }_{\mathbf{b}}$	${\mathrm{F}}_{\mathrm{b}}^{\mathrm{c}\mathrm{a}\mathrm{t}\mathrm{c}\mathrm{h}}/{\mathrm{F}}_{\mathrm{b}}$	dimensionless clutch catch bond rupture force	adjusted
${\beta }_{\mathbf{c}}$	${{\kappa }_{\mathrm{c}}\propto \beta }_{\mathrm{c}}$	clutch stiffness parameter	adjusted
${\mathbf{\omega }}^{\prime }$	$\omega /{\beta }_{\mathrm{c}}$	–	adjusted
${\mathrm{K}}^{\text{'}}$	$\mathrm{K}{\beta }_{\mathrm{c}}$	–	adjusted

^aParameter values are estimated in the supporting material.