Level of pathogen reduction achieved across various times of vaccination with different average host lifespans indicated by the different colors. Solid lines represent the level of pathogen reduction achieved for a given date of vaccine introduction. Dashed lines and the grey region beneath them represent the birthing season. Day 1 corresponds to the first day of the birthing season as well as the first possible day of vaccine introduction. The remaining parameters are: an average population size of 2000 individuals (), s = 3, R0,V = 1.5, R0,P = 2, 250 vaccines are distributed each year (NV = 250), individuals can disseminate vaccine for 21 days on average (γV and γg = 21−1), individuals remain infectious with the pathogen for 21 days on average (γP = 21−1), the transferable vaccine is groomed off individuals after 6 days on average (α = 1/15000, and the pathogen is non-virulent (ν = 0).