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. 2023 Sep 9;22:178. doi: 10.1186/s12934-023-02181-4

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© The Author(s) 2023

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated in a credit line to the data.

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Fig. 1 — Schematic representation of genetic organization of the GAD genes (i.e. regulator gadR, glutamate/GABA antiporter gadC, glutamate decarboxylase gadB, and two promoters P_gadR and P_gadCB) in lactococci: ∆ (blue) deletions observed in all L. cremoris species relative to L. lactis, ∆ (red) deletion or ▼ (red) insertion only observed in L. cremoris species with the cremoris phenotype, ▼ (grey) insertion site for the IS981 element in L. lactis S642. (not drawn to scale)