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. Author manuscript; available in PMC: 2023 Oct 2.
Published in final edited form as: Neurosci Biobehav Rev. 2022 Sep 8;141:104855. doi: 10.1016/j.neubiorev.2022.104855

Table 2c.

Pathway-specific manipulations in object exploration tests. AAV: adeno-associated virus or recombinant adeno-associated virus; ACC: anterior cingulate cortex; BLA: basolateral amygdala; CeA: central nucleus amygdala; C.N.E.: compatible with no important effect; CNO: clozapine-N-oxide; DA: dopamine; DG: dentate gyrus; dHPC: dorsal hippocampus; DLS: dorsolateral septum; h: hours; iCA1: intermediate CA1; IPN: interpeduncular nucleus; ISI: inter-sample interval; ITI: inter-trial interval; LC: locus coeruleus; LEC: lateral entorhinal cortex; Lenti: lentiviruses; mHb: medial habenula; min: minutes; mPFC: medial prefrontal cortex; MS: medial septum; NAc: nucleus accumbens; OVX: ovariectomy; pdCA1: posterior dorsal CA1; s: seconds; SuM: supramammillary nucleus; TH: tyrosine hydroxylase; vHPC: ventral hippocampus; VTA: ventral tegmental area; (+) excitation; (−) inhibition. *Two novel objects were replaced out of four objects. **The manipulated group was still able to show a preference for the novel object.

Animal Treatment Time of activation Encoding ITI Findings Reference
NOP test:
Lister Hooded
male rats
(−) pdCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into pdCA1
Days before
Daun02 4 μg/μl
4 min x4 1 h C.N.E. Barker et al., 2017
Lister Hooded
male rats
(−) iCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into iCA1
Days before
Daun02 4 μg/μl
5 min 1 h C.N.E.* Barker et al., 2017
Gad2-Cre
male mice
(−) LEC GABAergic terminals in dCA1
AAV2/9-Syn-FLEX-PSAM into LEC, cannulae in dCA1
Pre-sample
PSEM308 15 μM/0.5 μl
10 min x2 10 min Blunted** Basu et al., 2016
C57BL/6J
male mice
(+) selectively CeA neurons -< LC
AAV-CAG-Cre into LC + AAV-hSyn-DIO-hM3Ds into CeA
Pre-sample
CNO 1 mg/kg, i.p.
3-4 minx3 1 h Facilitated Paretkar & Dimitrov, 2018
Long Evans
male TH-Cre rats
(−) LC TH terminals in BLA
AAV2-hSyn-DIO-hM4Di into LC, cannulae in BLA
Pre-sample
CNO 3 μM/0.5 μl
10 min 3 h
24 h
C.N.E.
C.N.E.
Llorca-Torralba et al., 2019
Long Evans
male TH-Cre rats
(+) LC TH terminals in BLA
AAV2-hSyn-DIO-hM3Ds into LC, cannulae in BLA
Pre-sample
CNO 3 μM/0.5 μl
10 min 3 h
24 h
C.N.E.
C.N.E.
Llorca-Torralba et al., 2019
C57BL/6J mice
both sexes
(−) selectively subiculum excitatory neurons -< CA1
CAV2-Cre into CA1 + AAV2-DIO-hM4Di into subiculum
Pre-sample
CNO 1.5 mg/kg, i.p.
10 min 24 h C.N.E. Sun et al., 2019
C57BL/6J mice
either sex
(−) selectively CA1 neurons -< dorsal CA3
CAV2-Cre into CA3 + AAV2-DIO-hM4Di into CA1
Pre-sample
CNO 5.0 mg/kg, i.p.
10 min 24 h Impaired Lin et al., 2021
C57BL/6 mice
female OVX
(−) mFPC + dHPC excitatory neurons
AAV8-CaMKIIα-hM4Di into mPFC
AAV8-CaMKIIα-KORD into dHPC
Post-sample
CNO + SALB subdoses
reach 30s 24 h Impaired Tuscher et al., 2018
C57BL/6
male mice
(+) selectively vHPC neurons -< mPFC
AAV2-Cre into mPFC + AAV8-DIO-hSyn-hM3Dq into vHPC
Pre-test
CNO 3 mg/kg, i.p.
10 min x7 24 h C.N.E. Phillips et al., 2019
C57BL/6J
male mice
(−) MS neurons -< dHPC
AAV2-Cre into dHPC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
5 min 1 h C.N.E. Jin et al., 2020
C57BL/6J
male mice
(−) MS neurons -< ACC
AAV2-Cre into ACC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
5 min 1 h C.N.E. Jin et al., 2020
C57BL/6
male mice
(−) CA2/3 excitatory terminals in CA1
AAV5-CaMKIIα-eNpHR into CA2/3 & opto-fiber into CA1
During test trial 5 min x3 5 min Impaired Raam et al., 2017
C57BL/6
male mice
(−) CA2/3 excitatory terminals in pCA1
AAV5-CaMKIIα-eNpHR into CA2/3 & opto-fiber into pCA1
During test trial 5 min x3 5 min C.N.E. Raam et al., 2017
C57BL/6
male mice
(−) CA2/3 excitatory terminals in DLS
AAV5-CaMKIIα-eNpHR into CA2/3 & opto-fiber into DLS
During test trial 5 min x3 5 min C.N.E. Raam et al., 2017
Trpc4-Cre
male mice
(−) vCA1 excitatory terminals at NAc
AAV9-hSyn-DIO-eArchT into vCA1 & opto-fiber into NAc
During test trial
constant light
6 min x3 24 h C.N.E. Okuyama et al., 2016
DAT-Cre mice
both sexes
(+) VTA DA terminals in IPN
AAV2-Ef1a-DIO-ChR2 into VTA, opto-fiber in IPN
During test trial
30 Hz
5 minx2 24 h C.N.E. Molas et al., 2017
Chat-Cre mice
both sexes
(+) mHb choline-/glutamatergic terminals in IPN
AAV2-Ef1a-DIO-ChR2 into mHb, opto-fiber in IPN
During test trial
20 Hz
5 minx2 24 h Impaired Molas et al., 2017
Chat-Cre mice
both sexes
(−) mHb choline-/glutamatergic terminals in IPN
AAV2-Ef1a-DIO-NpHR3.0 into mHb, opto-fiber in IPN
During test trial
constant light
5 minx2 24 h Impaired Molas et al., 2017
NLP test:
Lister Hooded
male rats
(−) pdCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into pdCA1
Days before
Daun02 4 μg/μl
3 min 4 h C.N.E. Barker et al., 2017
Lister Hooded
male rats
(−) iCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into iCA1
Days before
Daun02 4 μg/μl
3 min 4 h C.N.E. Barker et al., 2017
TH-IRES-Cre mice
both sexes
(+) LC TH terminals in dHPC
AAV2/5-EF1a-DIO-ChR2 into LC + optic fiber in dHPC
During sample trial
20 Hz
5 min 24 h Facilitated Kempadoo et al., 2016
Rosa-LSL-ChR2
both sexes
(+) selectively subiculum neurons -< CA1
AAVrg-hSyn-Cre into CA1 + opto-fiber in subiculum
During sample trial
6 Hz
3 min 24 h Facilitated Sun et al., 2019
Vgat-Cre mice
both sexes
(+) SuM terminals in DG
AAV5-EF1α-DIO-ChR2 into SuM + optic fiber in DG
During sample trial
10 Hz
5 min 6 h C.N.E. Li et al., 2020a
Vgat-Cre mice
both sexes
(−) SuM terminals in DG
AAV5-EF1α-DIO-Arch3.0 into SuM + optic fiber in DG
During sample trial
constant light
5 min 6 h C.N.E. Li et al., 2020a
C57BL/6J mice
both sexes
(−) selectively subiculum excitatory neurons -< CA1
CAV2-Cre into CA1 + AAV2-DIO-hM4Di into subiculum
Pre-sample
CNO 1.5 mg/kg, i.p.
10 min 24 h Impaired Sun et al., 2019
C57BL/6J mice
either sex
(−) selectively CA1 neurons -< dorsal CA3
CAV2-Cre into CA3 + AAV2-DIO-hM4Di into CA1
Pre-sample
CNO 5.0 mg/kg, i.p.
10 min 24 h Impaired Lin et al., 2021
C57BL/6 mice
female OVX
(−) mFPC + dHPC excitatory neurons
AAV8-CaMKIIα-hM4Di into mPFC +
AAV8-CaMKIIα-KORD into dHPC
Post-sample
CNO + SALB subdoses
reach 30s 4 h Impaired Tuscher et al., 2018
Vgat-Cre mice
both sexes
(+) SuM terminals in DG
AAV5-EF1α-DIO-ChR2 into SuM + optic fiber in DG
During test trial
10 Hz
5 min 6 h Facilitated Li et al., 2020a
Vgat-Cre mice
both sexes
(+) SuM terminals in CA2
AAV5-EF1α-DIO-ChR2 into SuM + optic fiber in CA2
During test trial
10 Hz
5 min 6 h C.N.E. Li et al., 2020a
Vgat-Cre mice
both sexes
(−) SuM terminals in DG
AAV5-EF1α-DIO-Arch3.0 into SuM + optic fiber in DG
During test trial
constant light
5 min 6 h Impaired Li et al., 2020a
C57BL/6J
male mice
(−) MS neurons -< dHPC
AAV2-Cre into dHPC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
5 min 1 h Impaired Jin et al., 2020
C57BL/6J
male mice
(−) MS neurons -< ACC
AAV2-Cre into ACC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
5 min 1 h C.N.E. Jin et al., 2020
OiP test:
Lister Hooded
male rats
(−) pdCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into pdCA1
Days before
Daun02 4 μg/μl
5 min 1 h C.N.E. Barker et al., 2017
Lister Hooded
male rats
(−) iCA1 neurons -< mPFC
Lenti-lacZ into mPFC + cannule into iCA1
Days before
Daun02 4 μg/μl
5 min 1 h Impaired Barker et al., 2017
C57BL/6J
male mice
(−) MS neurons -< dHPC
AAV2-Cre into dHPC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
10 min 1 h Impaired Jin et al., 2020
C57BL/6J
male mice
(−) MS neurons -< ACC
AAV2-Cre into ACC + AAV9-DIO-hM4Di into MS
Pre-test
CNO 1.0 mg/kg, i.p.
10 min 1 h C.N.E. Jin et al., 2020