a Correlation between morphological and anatomical traits b Correlation between ionic contents and organic osmolytes SL: shoot length , RL: Root length, SFW: shoot fresh weight, RFW: root fresh weight, SDW: Shoot dry weight, RDW: root dry weight, S.CCA: stem cortical cell area, S.MxA: stem metaxylem area, S.PhA : stem phloem area, S.VBA: stem vascular bundle area, R.Eth: root epidermis thickness, R.CCA: root cortical cell area, R.MxA: root metaxylem area, R.PhA: root phloem area, PCA: pith cell area, R.ACA: root aerenchyma area, L.LTh: leaf lamina thickness, L.MTh: leaf midrib thickness, L.PhA: leaf phloem area, L.MXA: leaf metaxylem area, AB.SD: Abaxial stomatal density, Ab.SA: abaxial stomatal area, Ad.SD: adaxial stomatal density, Ad.SA: adaxial stomatal area, GB: Glycine betaine, PL: Proline, TSS: Total soluble sugars, TSP: total soluble proteins, F.AA: Free amino acids, S.Na: shoot sodium, R.Na: root sodium, S.Ca: shoot calcium, R.Ca: root calcium, S.K: shoot potassium, R.K: root potassium