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Determination of transcriptional start sites by high-resolution S1 protection assay. Maxam-Gilbert sequencing ladders (G+A and T+C reactions) were generated from the same 32P-labeled probe DNA fragments used in each S1 protection reaction. The positions of the S1-protected fragments are shown by the arrowheads, and the transcriptional initiation sites are assigned to the residues indicated by the bent arrows.
