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Extracellular activities of MnP (A) and LiP (B) as well as physiological concentrations of veratryl alcohol (C) and oxalic acid (D) during the time course experiment in manganese-free biobleaching cultures with EDTA-extracted pulp. The control culture not receiving organic acids was set at an initial pH of 6.0 (□). The glycolate (3 mM)-amended culture was set at an initial pH of 4.5 (▴). Error bars indicate standard deviations.
