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Phylogenetic placement of bacterial symbionts in O. crassitunicatus based on 16S rRNA sequences: maximum-likelihood trees of members of the γ-Proteobacteria (a), δ-Proteobacteria (b), and Spirochaeta (c). Symbionts of gutless oligochaetes are indicated by boldface type, and the O. crassitunicatus symbionts are enclosed in boxes. Bar = 10% estimated sequence divergence.
