| NO → [Pseudomonas] →N2O → [Pseudomonas] → N2 → [Azospirillum] → NH4+ → | (2) |
| [COMAMMOX; Nitrospira, Beijerinckiaceae] → NO2− → [COMAMMOX; Nitrospira, | |
| Beijerinckiaceae] → NO3− → [ANME] → NO2− → [ANME] →NO |
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