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. 2023 Nov 16;13:1251355. doi: 10.3389/fonc.2023.1251355

Table 2.

Metabolic effects on immune cells impact immune regulation.

Parasite infection or stimuli Treatment or model Metabolic effects in immune cells Metabolic effects at tissue and systemic level References
Secretion of IL-4 Promotion of OXPHOS and mitochondrial respiration in M2 macrophages surrounding the parasite (5)
Activation of M2 macrophages with IL-4 Bone marrow-derived macrophages isolated from femurs and tibias, stimulated with murine IL-4 Enhanced FAO promoted by STAT6 and PGC1β. Inhibition of proinflammatory cytokine production (5, 106)
IL-4R signaling in M2 macrophages Myeloid-cell-specific IL-4Rα-deficient mice (Il4ra fl/− Lyz2-cre) and Retnla −/− mice M2 macrophages activated by IL-4R signaling express IGF-1, RELMα and Arg-1. RELMα and Arg-1 enhance wound healing. (104, 107)
Stimulation with IL-4 CD36 expression induced by IL-4 in RAW 264.7 cells Induction of CD36 expression and lysosomal function with endocytosis of LDL and VLDL (108)
Brugia malayi Human monocyte-derived dendritic cells exposed to B. malayi microfilaria from infected jirds Downregulation of components of the mTOR signaling pathway in microfilaria-induced DC. Inhibition of mTOR and its regulatory proteins phosphorylation which are vital for protein synthesis in DC. Increased autophagy (109)
Nippostrongylus brasiliensis RIP2-Opa1KO mice with pancreatic β cell Opa1 deficiency, infected trough subcutaneous inoculation of third stage N. brasiliencis larvae Increase in WAT eosinophils and M2 macrophages with increased expression of M2 markers (YM1 and Arg-1). Increase in body insulin sensitivity and glucose tolerance, diminished hepatic steatosis (41, 110)
Heligmosomoides polygyrus C57BL/6 mice fed with high-fat diet and infected with 200 third stage H. polygyrus larvae Decreased weight gain, increase in glucose tolerance and WAT beiging. Increase in WAT M2 gene expression and M2 markers (41, 111)
Fasciola hepatica Bone marrow-derived macrophages and peritoneal macrophages from C57BL/6 mice, stimulated with synthetic FhHDM-1 FhDHM-1 reprograms macrophages by inducing OXPHOS and elevation of glutaminolysis, resulting in inhibition of pro-inflammatory cytokines (TNF and IL-6) independent of M2 polarization. Inhibition of lysosomal vATPSase. (112)
Schistosoma mansoni Arg1(-/flox); LysMcre mice and Arg1(flox/flox);Tie2cre mice Macrophage expression of Arg-1, triggered by S. mansoni infection, downregulates inflammation and T proliferation by depleting arginine concentrations. Arg-1 expressing macrophages have an anti-fibrotic activity during Th2 response, and are important mediators of immune modulation of chronic schistosomiasis (113)
Heligmosomoides polygyrus A 2B AR −/− BL/6 mice infected with third stage H. polygyrus larvae trough oral administration Adenosine initiates a helminth-induced type 2 response through interaction with the A2B adenosine receptor. Upregulation of IL-33 and the subsequent activation of ILC2 cells (104, 114)

Acronyms, OXPHOS, oxidative phosphorylation; Arg-1, arginase 1; WAT, white adipose tissue; RELMα, resistin-like molecule alpha. FhHDM-1, Fasciola hepatica helminth defense molecule; DC, dendritic cell. mTOR, mammalian target of rapamycin.

—, Not determined.