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. 2023 Aug;15(4):574–584. doi: 10.18502/ijm.v15i4.13512

Table 2.

Bacterial diversity from various sampling methods on holy handwritten Qurans.

SDA TSA SDA TSA Frequency


S1 S2
S1 Staphylococcus warneri Staphylococcus warneri
Bacillus halotolerans, Bacillus atrophaeus **
Paenibacillus polymyxa
Cellulosimic robium aquatile
Bacillus zhangzhouensis
Acinetobacter junii
Bacillus atrophaeus Bacillus atrophaeus
Cellulosimic robium aquatile *
16 (53.3%)
S3 Bacillus atrophaeus ** Terribacillus goriensis
Bacillus atrophaeus
Bacillus mobilis
Bacillus atrophaeus Bacillus paramycoides
Bacillus atrophaeus
9 (30%)
S2 Staphylococcus epidermidis Staphylococcus warneri Staphylococcus warneri Pantoea eucrina 5 (16.7%)
Total 5 (16.67%) 12 (40%) 4 (13.3%) 9 (30%) 30 (100%)

S1: Sample 1 (parchment). S2: Sample 2 (Manuscript belong to the tenth century). S3: Sample 3 (Manuscript belongs to the tenth century).

*Four isolates of Cellulosimicrobium aquatile were detected.

**Three isolates of Bacillus atrophaeus were detected.