Design of CRISPR-Cas antifungals. (A, B) CRISPR-Cas
antifungal
formulation (A) with essential gene targeting alone and (B) with essential
and defensive gene cotargeting. Targeting a single essential gene
locus alone is not effective to eliminate cell survival due to active
DNA repair machinery. Cotargeting parts of the DNA repair response
reduces the cell’s capability to repair legions at essential
and defensive sites, increasing lethality via essential gene knockout
and DSB-induced apoptosis. (C, D) Heat maps show the effectiveness
of (C) single essential or defensive gene targeting and (D) essential
and defensive gene cotargeting. Each heat map represents serial dilution
(1×, 10×, 100×, and 500×) of cells that were treated
with different CRISPR-Cas antifungals, spotted on galactose plates
(Cas9 “on”, test) and glucose plates (Cas9 “off”,
control), and incubated for 48 h at 28 °C. Noninduced strains
grown on glucose plates are indicated in red, while induced Cas9 strains
grown on galactose pads are indicated in purple. The top 17 strains
chosen for further characterization are marked with a star. Larger
colony size is represented by darker coloration. Note that panels
B and D show a representative list of characterized strains; the complete
list can be found in Figure S1. (E) Gene
cotargeting interaction analysis.