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. 2023 Nov 22;9(12):1128. doi: 10.3390/jof9121128
1. Pileus with blackish-brown, reddish-brown, yellowish-brown, brown or golden-tawny scales 2
1. Pileus with black, greyish-black, grey or dirty white scales 17
2. Pileus with golden-tawny to golden-orange scales; basidiospores smaller
measuring 6–8 × 5.5–7 μm
S. mirandus
2. Pileus with blackish-brown, reddish-brown, brown or yellowish-brown scales; basidiospores larger, more than 7 μm in length 3
3. Pileus smaller (up to 4 cm), with yellowish-brown scales S. minor
3. Pileus larger (up to 12 cm), with blackish-brown, reddish-brown or brown scales 4
4. Pileus with reddish-brown or brown scales 5
4. Pileus with blackish-brown scales 9
5. Stipe with dirty white to light reddish-brown thin flosses; hymenophoral pores smaller, 0.05–0.1 cm diameter S. glabellus
5. Stipe with reddish-brown, brown, dark brown to blackish-brown fluffy flosses; hymenophoral pores larger, 0.1–0.3 cm diameter 6
6. Pileus with reddish-brown to vinaceous brown on the lower part to dark brown to blackish-brown on the upper part, crowded, more or less erect pyramidal scales; stipe with dark brown to blackish-brown fluffy flosses S. atrosquamosus
6. Pileus with reddish-brown or brown, scattered, erect conical scales; stipe with concolorous fluffy flosses and conical scales 7
7. Hymenophoral pores smaller, 0.05–0.1 cm diameter; stipe with light grayish-brown to chocolate fluff on the upper part and reddish-brown to chocolate fluff on the lower, basidiospores larger measuring 8–10 × 7–8 μm S. rubrobrunneus
7. Hymenophoral pores larger, 0.1–0.3 cm diameter; stipe with brown or reddish fluffy flosses or scales, basidiospores smaller, less than 9 μm in length 8
8. Scales on pileus brown, without reddish tinge; stipe surface reticulate with shallow and elongate meshes on the upper part, covered with clustered tomentose scales on the lower part; basidiospores smaller measuring 7–8 × 6.5–7 μm S. sculptus
8. Scales on pileus with reddish tinge; stipe with concolorous thick fluffy flosses and erect conical scales; basidiospores larger measuring 7.5–9 × 6.5–8 μm S. brunneolepidotus
9. Pileus with more or less erect conical to pyramidal scales 10
9. Pileus with patch-like to appressed scales or flosses 15
10. Basidiospores with complete reticula; hymenophoral pores larger, 0.05–0.5 cm diameter 11
10. Basidiospores with incomplete reticula; hymenophoral pores smaller, 0.05–0.1 cm diameter S. conicus
11. Hymenophoral pores larger, 0.1–0.5 cm diameter; cystidia thick-walled (up to 3 μm); tropical distribution S. pachycystidiatus
11. Hymenophoral pores smaller, 0.05–0.15 cm diameter; cystidia thin-walled, less than 1 μm; temperate, tropical or subtropical distribution 12
12. Stipe with blackish-brown thin fluffy flosses 13
12. Stipe with sooty-brown to blackish, grey to dirty white or greyish-black flosses 14
13. Pileus with blackish-brown to vinaceous, soft erect conical scales; basidiospores with larger meshes (2–3.5 μm diameter) S. mollis
13. Pileus with black-brown, more or less erect pyramidal scales; basidiospores with smaller meshes (1–2 μm diameter) S. parvirimosus
14. Pileus with sooty-brown to blackish scales; stipe without an annulus, with cottony or woolly, sooty-brown to blackish floccules S. echinocephalus
14. Pileus with dirty white to light blackish-brown scales; stipe with an annulus, with grey to dirty white on the upper part and greyish-black flosses on the lower part S. microreticulatus
15. Hymenophoral pores larger, 0.1–0.2 mm diameter; basidiospores larger measuring 11.5–14 × 9.5–11 μm, with larger meshes (2–4 μm diameter) S. alpinus
15. Hymenophoral pores smaller, 0.05–0.1 mm diameter; basidiospores smaller, less than 11 μm in length, with smaller meshes (1–3 μm diameter) 16
16. Pileus with blackish-brown at apex and light brown to dirty white at base, scales or flosses; stipe with an annulus, with greyish-white fluffy flosses on the upper part and dark blackish-brown on the lower arranged in spiral; basidiospores larger measuring 9–11 × 7–8.5 μm; subtropical to temperate distribution S. cingulatus
16. Pileus with blackish-brown to dark chocolate scales or flosses; stipe without an annulus, with dirty white thin flosses evenly distributed; basidiospores smaller measuring 7–9 × 6–7 μm; tropical distribution S. albidus
17. Pileus with black scales 18
17. Pileus with greyish-black, grey to dirty white scales 24
18. Pileus with more or less erect conical to pyramidal scales 19
18. Pileus with patch-like to appressed scales or flosses 23
19. Basidiospores with complete reticula 20
19. Basidiospores with incomplete reticula 22
20. Pileus larger (up to 9.7 cm); stipe with thick flosses arranged in spiral S. douformis
20. Pileus smaller (up to 5 cm); stipe with thin flosses evenly distributed 21
21. Pileus with charcoal black, smaller, more or less erect pyramidal scales; basidiospores smaller measuring 8.5–10 × 6.5–8(–9) μm; subtropical distribution S. anthracinus
21. Pileus with black erect conical to pyramidal scales; basidiospores larger measuring 9–11.5 × 8–10 μm; tropical distribution S. hainanensis
22. Pileus with soft, more or less erect conical to pyramidal scales; stipe without an annular zone, entirely with black granular scales; echinate basidiospores with confluent tubercles and irregular incomplete reticulations S. calidus
22. Pileus with hard, more or less erect conical scales; stipe with an annular zone at apex, with black minutely conical scales and fluffy flosses; basidiospores with semireticulate ornamentation S. giganteus
23. Pileus larger (up to 6 cm); stipe with patch-like to appressed, blackish-brown scales or reticula; tropical distribution S. baozhengii
23. Pileus smaller (up to 4.5 cm); stipe with black tomentose scales; subtropical distribution S. huangshanensis
24. Pileus with more or less erect conical to pyramidal scales 25
24. Pileus with patch-like to appressed scales or flosses 29
25. Basidiospores with complete reticula 26
25. Basidiospores with incomplete reticula 28
26. Stipe without an annular zone; tropical to subtropical distribution S. montosus
26. Stipe with an annular zone; subtropical to temperate distribution 27
27. Pileus larger (up to 11 cm), with dirty white, small, erect conical scales and greyish-black apex; hymenophoral pores larger, 0.1–0.2 cm diameter S. pteroreticulosporus
27. Pileus smaller (up to 7 cm), with greyish-black, more or less erect pyramidal scales; hymenophoral pores smaller, 0.05–0.1 cm diameter S. pinophilus
28. Pileus larger (up to 12 cm); stipe with greyish-white flosses on the upper part and black on the lower; echinate basidiospores with irregular short ribs; subtropical to temperate distribution S. densisquamosus
28. Pileus smaller (up to 6 cm); stipe with black scattered granular scales; semireticulate basidiospores S. velutinus
29. Basidiospores with complete reticula 30
29. Basidiospores with incomplete reticula 31
30. Pileus larger (up to 15 cm); stipe with an annulus, with grey to dirty white flosses on the upper part and greyish-black on the lower; basidiospores with larger meshes (2–4 μm diameter); subtropical distribution S. glabriceps
30. Pileus smaller (up to 9 cm); stipe without an annular zone, entirely with whitish thick fluffy flosses; basidiospores with smaller meshes (1–2 μm diameter); tropical and subtropical distribution S. latirimosus
31. Stipe with an annular zone; basidiospores smaller measuring 7–9 × 6.5–8.5 μm; tropical to subtropical distribution S. seminudus
31. Stipe without an annular zone; basidiospores larger measuring 8–10 × 7.5–8.5 μm; subtropical distribution S. subnudus