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. 2023 Dec 15;14:1303667. doi: 10.3389/fpls.2023.1303667

Table 3.

Mechanism of WRKY on metabolism of flavonoids.

Species WRKY name Subgroup Regulation of target gene Regulation of biological Target gene Function References
Arabidopsis thaliana AtWRKY41 3 / / Negative regulation of anthocyanin biosynthesis (Duan et al., 2018)
Malus crabapple McWRKY71 2c + + McANR Enhance proanthocyanidin biosynthesis (Zhang et al., 2022b)
Malus domestica MdWRKY75 2c + + MdMYB1 Enhance anthocyanin accumulation (Su et al., 2022b)
Malus domestica MdWRKY71-L 2c + + MdUFGT and MdMYB1 Enhance anthocyanin accumulation (Su et al., 2022a)
Malus domestica MdWRKY40 2a + + MdUFGT Promote wounding-induced anthocyanin biosynthesis (An et al., 2019)
Malus domestica MdWRKY11 2d + + MdUFGT Enhance anthocyanin accumulation (Liu et al., 2019a)
Malus domestica MdWRKY41 3 MdANR, MdUFGT and MdMYB12 Negatively regulates anthocyanin and PA biosynthesis (Mao et al., 2021)
Malus domestica MdWRKY72 2b + + MdHY5 and MdMYB1 Enhance anthocyanin accumulation (Hu et al., 2020)
Malus domestica MdWRKY1 2d / + / Enhance anthocyanin accumulation (Ma et al., 2021a)
Vitis vinifera VvWRKY26 1 + + VvCHI Improved activation efficiency and flavonoid accumulation (Amato et al., 2019)
Vitis vinifera VqWRKY56 2b + + VvCHS3, VvLAR1, and VvANR Promote proanthocyanidin biosynthesis and increase resistance to powdery mildew (Wang et al., 2023d)
Pyrus L. PpWRKY44 1 + + PpMYB10 Stimulating anthocyanins (Alabd et al., 2022)
Pyrus bretschneideri PbWRKY75 2c + + PbDFR, PbUFGT, and PbMYB10b Promote anthocyanin synthesis (Cong et al., 2021)
Pyrus L. PyWRKY26 1 + + PyMYB114 Promote anthocyanin synthesis (Li et al., 2020a)

ANR, anthocyanin reductase; ANS, anthocyanidin synthase; CHI, chalcone isomerase; CHS, chalcone synthase; DFR, dihydroflavonol 4-reductase; F3H, flavanone 3-hydroxylase; HY, ELONGATED HYPOCOTYL; LAR, leucine anthocyanin reductase; MYB, myb avian myeloblastosis viral oncogene homolog; UFGT, flavonoid 3-O-glycosyl-transferase. The ‘+’ represents positive regulation, the ‘-’ represents negative regulation, and the ‘/’ represents unknown.