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. 2023 Dec 21;14:1295632. doi: 10.3389/fphys.2023.1295632

TABLE 1.

Genes used in generating transgenic rodent lines for studying respiratory rhythm generation.

Gene Line Type Reference Notes
Atp1a2 Atp1a2−/− CKO Ikeda et al. (2004) Brain-specific due to gene expression pattern
Atoh1 Atho1 Cre∕− ; Tau mgfp−nLacZ CKO Huang et al. (2012) Expression in hindbrain during development
Atho1 Phox2bCKO CKO Expression preferentially in retrotrapezoid nucleus
Atoh1 HoxA4CKO CKO Expression in caudally derived Atho1 lineages
chx10 Chx10:LNL:DTA CKO Crone et al. (2012) Ablation of V2a neurons by diphtheria toxin A-chain (DTA)
Dbx1 Dbx1+/CreERT2; Rosa26 tdTomato CKI Wang et al. (2014) Identification of Dbx1 neurons
GluR3 GluR3−/− KO Steenland et al. (2008) Constitutive knockout
Jmjd3 Jmjd3−/− KO Burgold et al. (2012) Neurogenic factor required for maintenance of the preBötC
Jmjd3−/−; Jmjd3 mut 21 KO Point mutation A1388H
Jmjd3−/−; Jmjd3 mut 65 KO Point mutation A1388H
Jmjd3−/−; Rosa26 Jmjd3 KO/CKI Conditional rescue of Jmjd3
Rosa26 flSTOPJmjd3 CKI Conditional expression of Jmjd3
Lbx1 Lbx1 GFP∕GFP KO Pagliardini et al. (2008) Expression during development of medulla and spinal cord
MAO-A Tg8 KO Bou-Flores et al. (2000) Deletion of gene. Displays excess of 5-HT.
Mafb Mafb−/− KO Blanchi et al. (2003) Constitutive knockout. Differentiation of hematopoietic system, podocytes, and specification of rhombomeres in the early hindbrain
Mecp2 B6.129P2(C)-MeCP2 tm1−1Bird KO Mironov et al. (2009) Constitutive knockout. Transduction of neurons in organotypic slices with a Ca2+sensor (D3cpv) via AAV.
Rnx Rnx−/− KO Shirasawa et al. (2000) Orphan homeobox gene expressed in the developing dorsal and ventral region of the medulla
Sst Sprague-Dawley KI Tan et al. (2008) Transduction with AAV2-pA-Sst-alstR-IRES-EGFP or AAV2-pA-Sst-EGFP into the preBötC is performed to express the allatostatin receptor and EGFP (enhanced green fluorescent protein) in Sst neurons. Alternatively, only EGFP is expressed in Sst neurons using AAV2-pA-Sst-EGFP.
STIM1 STIM1−/− KO Berry et al. (2003) Highly expressed in the central nervous system and placenta
Thy1.2 Thy1.2-EYFP KI Winter et al. (2007) Expression of EYFP (enhanced yellow fluorescent protein) in the ventral respiratory column, preBötC, and the rostral ventral respiratory group neurons
Thy1.2-HcRed KI Expression of HcRed in the ventral respiratory column, preBötC, and the rostral ventral respiratory group neurons
Tshz3 Tshz3+/lacZ KO Caubit et al. (2010) Transcription factor expressed in the brainstem
Vglut2 Vglut2 f/f;PCre CKO Wallén-Mackenzie et al. (2006) First model showing complete uncompensated functional impairment of the respiratory CPG.
Vglut2, Dbx1, and Sst Dbx1-tdTomato CKI Koizumi et al. (2016) Expression of tdTomato in Dbx1-neurons
Dbx1-tdTomato-Arch-GFP CKI Expression of tdTomato, archaerhodopsin (Arch), and GFP in Dbx1-neurons
Sst-tdTomato CKI Expression of tdTomato in Sst-expressing neurons
SST-tdTomato-Arch-GFP CKI Expression of tdTomato, Arch, and GFP in Sst-expressing neurons
VgluT2-tdTomato CKI Expression of tdTomato in VgluT2-expressing neurons
VgluT2-tdTomato-Arch-GFP CKI Expression of tdTomato, Arch, and GFP in VgluT2-expressing neurons
VgluT2-tdTomato-Gad67-GFP CKI Expression of tdTomato in VgluT2-expressing neurons and expression of GFP in Gad67-expressing neurons
ZFHX Zfhx4 PB/+ KO Zhang et al. (2021) Transcriptional factor expressed in the brainstem

Unless stated, all transgenic lines are from mice. Transgenic rats. Viral vectors. Atoh1/Math1 (Atonal BHLH, Transcription Factor 1), Atp1a2 (ATPase, Na+/K + Transporting Subunit Alpha 2), Chx10 (Visual system homeobox 2), Dbx1 (Developing Brain Homeobox 1), Gad67 (Glutamate Decarboxylase 67 KDa, Isoform), GluR3 (Glutamate Ionotropic Receptor AMPA, Type Subunit 3), Jmjd3 (Jumonji domain-containing protein-3), Lbx1 (Ladybird Homeobox 1), Mafb (MAF BZIP, Transcription Factor B), Maoa (Monoamine oxidase A), Mecp2 (Methyl-CpG, Binding Protein 2), Rnx (Respiratory Neuron Homeobox), Sst (Somatostatin), Stim1 (Stromal Interaction Molecule 1), Thy1 (Thy-1, cell surface antigen), Tshz3 (Teashirt Zinc Finger Homeobox 3), Vglut2 (Vesicular glutamate transporter 2), and Zfhx3 (Zinc Finger Homeobox 3).

KO, knockout; KI, knock-in; CKO, conditional knockout; CKI, conditional knock-in.