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Kinetics of β-Gal expression by in vitro-grown (A) and macrophage-grown (B) L. pneumophila harboring a ppa promoter (pPPZ) fused to the promoterless lacZ gene of pEU730. Data are expressed as the mean β-Gal activity for triplicate samples (34). L. pneumophila harboring the vector pEU730 did not show any detectable β-Gal activity (data not shown).
