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. 2024 Jan 10;625(7995):535–539. doi: 10.1038/s41586-023-06900-0

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© The Author(s) 2024

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 1 — a–c, The location of Southern China, Guangxi ZAR province and the city of Nanning (a), with the location of the Chongzuo study area marked by a large box (b) and the Bubing Basin study area marked by a smaller box (c). b, The location of the 16 cave sites analysed in the Chongzuo study area. c, The location of the six caves analysed in the Bubing Basin study area including both G. blacki-bearing and non-G. blacki-bearing caves from both regions.