Parameter	Meaning
$v\left[t\right],\theta h\left[t\right],p\left[t\right]$	Observed velocity, head curvature, and pumping rate
$n\left[t\right]$	Modeled neural activity.
$\mathrm{R}\mathrm{e}\mathrm{c}\mathrm{t}\left(c_{vT},v\left[t\right]\right)$	Locomotion direction rectification term with different values based on forwards versus reverse movement.
$c_{vT}$	Locomotion direction rectification parameter.
$c_v,c_{\theta h},c_p$	Velocity, head curvature, and feeding parameters.
$s$	Exponentially weighted moving average (EWMA) timescale parameter.
$b$	Baseline activity parameter.
$n\left[0\right]$	Initial condition parameter.
${\sigma }_{noise}$	White noise parameter.
${\sigma }_{\mathit{SE}}$	Autocorrelative residual parameter.
$\ell$	Autocorrelative residual timescale parameter.
$𝒢𝒫$	Gaussian process.
$K_{GN},K_{SE}$	Gaussian process kernels.