Full annual cycle of Greenland white-fronted geese. Sub-season designations used in analyses (a) included late winter, spring migration from wintering to Icelandic staging areas, first half of spring staging, second half of spring staging, spring migration from staging to Greenlandic breeding areas, early breeding, breeding, late breeding, autumn migration from breeding to Icelandic staging areas, first half of autumn staging, second half of autumn staging, autumn migration from staging to wintering areas, early winter and winter. Map (b) depicts GPS locations of all tracked geese from the study colour-coded by season (green = winter, dark blue = spring, orange = breeding, light blue = autumn).