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. 2024 Mar 4;18:1323555. doi: 10.3389/fnins.2024.1323555

Table 2.

The effect and mechanisms of acupuncture on cognitive disorders in neurodegenerative diseases.

Author Animal model/participant Acupoint Acupuncture type Acupuncture session Behavioral test Outcomes
Glutamatergic system
He et al. (2022) METH withdrawal mice model of spatial memory impairment GV29, GV20 EA, 200 Hz with 1–2 mA, 30 min/day 14 days MWM: time in probe test↓; time in OLM test↑ dCA1: extracellular Glu↓, c-Fos-positive neurons↓; C3-positive astrocytes↓, reactive astrocytes↓, C3 protein↓, GLT-1-positive astrocytes↑, GLT-1 protein↑;
Schaffer projections from dCA3 to dCA1: number of c-Fos-positive presynaptic neurons#, VGLUT-1-positive staining#
Hou et al. (2022) SAMP8 mice model of AD GV20, ST36 EA,1 mA, 10 Hz, 30 min/day 14 days Y Maze Test: correct spontaneous alternation rate↑; MWM test: movement paths↓, escape latency↓, percentage of time spent in target quadrant↑, number of platform crossings↑ CSF: OxA↓, Glu↓;
hippocampus: Nissl bodies (CA1, CA3)↑; OxA protein (hippocampus, lateral hypothalamus)↓, cAMP↑, pPKA/PKA↑, pCREB/CREB↑, GluN1↑, GluN2A↑, GluA2↑, SYP↑, PSD95 ↑, number of synapse↑, synaptic density↑
Lin et al. (2018) APP/PS1 transgenic mice model of AD GV20 EA, 100 Hz, 1/20 mA, 30 min/day 6 days/week, 4 weeks Escape latency↓, number of mice crossing the platform↑
Step-down avoidance test: latency↑, number of errors↓
Left hippocampus: NAA/Cr↑, Glu/Cr↑;
hippocampus, CA1 and CA3 areas: surviving neurons↑;
hippocampus: BDNF↑, TrkB↑
GABAergic system
Li et al. (2023) 5xFAD mice model of AD GV20, GV14 EA, 2 Hz, 1 mA, 30 min/day 5 days MWM test: escape latency↓
NOR task: time↑
Hippocampus: gephyrin protein↑, synapsin 1#, PSD95#, GAD65↑, SST↑, PV#;
dorsal CA1: CR↑ cell↑, sIPSCs frequency↑, sEPSCs frequency↑, sEPSCs area↑, sEPSCs amplitude#, charge ratio of sEPSC/sIPSC↓, theta oscillation↑, gamma oscillation↑;
dorsal hippocampus: SST+ cell↑
Serotonergic system
Xiao et al. (2023) D-galactose induced rat model of AD GV20, ST36 EA, 50 Hz, 1 mA, 20 min/day once a day, 8 weeks MWM test: escape latency↓, exploration time↑ colon: 5-HT levels↓, Trp↑
hippocampus: 5-HT levels↑, Trp↑, 5-HT6R↓, JNK↓, p-JUNK↓, c-JUN↓, p-c-Jun↓
Cholinergic system
Li et al. (2022) 5xFAD mice model of AD GV 20, GV 14 EA, 2/20 Hz, 1 mA, 30 min/day 5 days/week, 4 weeks OFT#: traveled distance, average speed, time spent in the central part;
NOR#: object exploration;
Novel location recognition: location exploration↑;
Location discrimination task: Must touch training trials#, PIT accuracy#, LDR trials↓
MS/VDB: NAA/Cr↑, Cho/Cr↑; AChE↓, VAChT↑, Aβ fraction ratio↓#
DG: NAA/Cr↑, Cho/Cr↑; DG, MOD of AChE#, MOD of VAChT#, ChAT↑, Aβ fraction ratio↓, M1+ cell↑
hippocampus: DCX+ cell↑, Neuro-D1+ cell↑
Lee et al. (2014) Scopolamine induced rat model of memory impairment GV20 MA - MWM test: escape latency↓ Hippocampus: ChAT↑, BDNF↑, CREB↑, CHT1↑, VAChT↑, BDNF mRNA↑, CREB mRNA↑
Lee et al. (2012) Corticosterone induced rat model of spatial cognitive impairment HT7/TE5 MA - MWM test: escape latency↓;
OFT: line crossing number#
Hippocampus: cholinergic neurons↑, BDNF mRNA↑, CREB mRNA↑
hippocampal CA1 area: ChAT-immunopositive neuron↑, AchE-immunopositive fibers↑;
hippocampal CA3 area: AchE-immunopositive fibers↑;
Medial septum: ChAT-immunopositive neuron numbers↑
Glial cells
Xie et al. (2021) Aβ1-42 induced rat model of AD GV20 EA, 10 mA, 20 Hz, 30 min/day 6 times/week, 3 weeks MWM test: escape latency↓, swimming speed↑, swimming distance↓, probe times of trails↑ CA1: activation of astrocytes↓, Iba1+ iNOS+ cell↓, Iba1+ Arg1+ cell↑, GFAP+ IL-1β+ cell↓, GFAP+ TNF-α+ cell↓, GFAP+ IL-6 cell↓, Iba1+ IL-4+ cell↑, Iba1+ IL-10+ cell↑, GFAP+ IL-4+ cell↑, GFAP+ IL-10 cell↑, Iba1+ p65+ cell↓, GFAP+ p65+ cell↓, Iba1+ STAT6+ cell↑, STAT6 mRNA expression↑;
DG: activation of astrocytes↓, Iba1+ iNOS+ cell↓, Iba1+ Arg1+ cell↑, IL-1β-#, TNF-α↓#, Iba1+ IL-6+ cell#, GFAP+ IL-1β+ cell↓, GFAP+ TNF-α+ cell↓, GFAP+ IL-6+ cell↓, Iba1+ IL-4+ cell↑, Iba1+ IL-10+ cell↑, GFAP+ IL-4+ cell↑, GFAP+ IL-10 cell↑, GFAP+ p65+ cell↓, Iba1+ STAT6+ cell↑, STAT6 mRNA expression↑
Li M. et al. (2020) SAMP8 mice model of AD GV20, GV29 EA, 2 Hz, 0.1 mA, 15 min/day once a day, 15 days MWM test: escape latency↓, platform crossing number↑ Hippocampus: TREM2 protein↑
Wang et al. (2020) SAMP8 mice model of AD GV29, LI20 EA, 15 Hz, 1.5 mA, 10 min/day 6 days/week, 4 weeks MWM test: escape latency↓, path length of place
navigation trial↓, swimming time↑
Hippocampus: Aβ protein↓, hyperphosphorylated tau proteins↓, P-tau/tau↓#, integrated density of Iba1+ cell↓, activated microglia area↓;
CA1: SYP expression content↑
Cai et al. (2019) 5xFAD mice model of AD KI3 EA, 1 mA, 2 Hz, 15 min/day 3 times/week, 2 weeks NOR test: percentage of exploration time of the novel object↑;
Y maze test: spontaneous alternations↑
Frontal cortex: mean glucose level↑;
prefrontal cortex: CD11b↓, GFAP↓, COX2↓, HO-1↓, transferrin↓, Bax↓, LTP↑, synaptophysin, PSD-95↑, synaptic structures↑, APP↓, APOE↓#, BACE↓#, Aβ1–42 levels↓, CD68+ cell/Aβ plaque↓;
cortex: mean glucose level↑;
hippocampus: mean glucose level#, LTP↑;
hypothalamus: mean glucose level↑
Tang et al. (2019) Aβ1-42 induced rat model of AD GV20, BL23 MA, 20 min/day once a day, 8 weeks MWM test: escape latency↓, platform crossing number↑ Hippocampal dentate gyrus: neuronal edema↓, astrocytes edema↓
Wang et al. (2014) APP/PS1 transgenic mice model of AD GV20 EA, 2/15 Hz, 1 mA, 30 min/day 5 days/week, 4 weeks MWM test: escape latency↓, swimming speed#, time spent in the target quadrant↑ GFAP↓, NDRG2↓
Han et al. (2018) LPS-induced mice model of cognitive impairment GV20 EA, 2/100 Hz, 4 mA, 30 min/day 7 days MWM test: escape latency↓, platform crossing number↑ Hippocampus: α7nAChR positive neurons↑, ACh content↑, ChAT↑, AChE activities↓, MDA↓, H2O2↓, CAT activity↑, GSH content↑, IL-1β↓, IL-6↓, TNF-α↓

#p > 0.05; 5 × FAD, five familial Alzheimer’s disease; α7nAChR, α7 nicotinic acetylcholine receptors; Aβ, amyloid-β; ACh, acetylcholine; AD, Alzheimer’s disease; APOE, apolipoprotein E; APP/PS1, amyloid precursor protein/presenili1; Arg1, M2 marker Arginase 1; BACE, β-site amyloid precursor protein cleaving enzyme 1; Bax, Bcl-2-associated X; BDNF, brain-derived neurotrophic factor; C3, compliment component 3; cAMP, cyclic adenosine monophosphate; CAT, catalase; CD11b, cluster of differentiation 11b; Cho, Choline; Cr, creatine; CREB, cAMP response element-binding protein; CSF, cerebrospinal fluid; COX2, cyclooxygenase 2; dCA1, dorsal CA1; EA, electricacupuncture; GABA, γ-aminobutyric acid; GFAP, glial fibrillary acidic protein; GLT-1, glutamate transporter 1; Glu, glutamine; GSH, glutathione; H2O2, hydrogen peroxide; IL, interleukin; iNOS, inducible nitric oxide synthase inos; LDT, Location Discrimination Task; LPS, lipopolysaccharide; LTP, long-term potentiation; MA, manual acupuncture; MDA, malondialdehyde; MEPs, Motor-Evoked Potentials; MFB, medial forebrain bundle; mGluR3, metabotropic glutamate receptor 3; MRI, Magnetic Resonance Imaging; MS/VDB-DG, medial septal and vertical limb of the diagonal band and dentate gyrus; mTOR, mammalian target of rapamycin; MPTP, 1-methyl-4-phenyl-1,2,3,6-tetrahydropyridine; MWM, Morris water maze; NAA, N-acetylaspartate; NG2, nerve/glial antigen 2; NDRG2, N-myc downstream-regulated gene 2; NeuN, neuron-specific nuclear protein; NLR, Novel Location Recognition; NOR, novel object recognition; OFT, Open Field Test; OLM, object location memory; ORT, Object Recognition Test; OxA, Orexin A; pCREB, phosphorylated CREB; PIT, punish incorrect training; PKA, cAMP response element-binding protein; PSD95, postsynaptic density protein 95; PV, parvalbumin; SAMP8, senescence-accelerated mice prone 8; sEPSCs, spontaneous excitatory postsynaptic currents; SST, somatostatin; STAT6, signal transducer and activator of transcription 6; STN, subthalamic nucleus; SYP, synaptophysin; TREM2, triggering receptors expressed on myeloid cells 2; Trp, tryptophan; TrkB, Tyrosine Kinase receptor B; VAChT, vesicular acetylcholine transporter; VGLUT-1, vesicular Glu transporter 1.