Table 1. Evolutionary parameters appearing in the model system.

Symbol	Description	Units
${\displaystyle N_M}$	Carrying capacity of the mature population
${\displaystyle t_M}$	Age when the cell population reaches mature size	years
${\displaystyle N_H}$	Population size at homeostatic divisions(start of the mixed-growth phase)
${\displaystyle \gamma }$	Symmetric division rates in early developmental phase	/year
${\displaystyle \rho }$	Symmetric division rate in homeostatic state	/year
${\displaystyle \mathrm{\Phi }}$	Asymmetric division rate in homeostatic state	/year
${\displaystyle \mu }$	Mutation rate	/division/daughter