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. 2024 Mar 6;15:1289492. doi: 10.3389/fimmu.2024.1289492

Table 2.

Compilation of studies regarding Sjögren’s syndrome and DNA methylation in cells.

Country
Reference
Study design for DNA methylation Cell types Main outcomes for DNA methylation
Norway
BrækkeNorheim K
et al.
(76)
Methylation analysis of patients in pSS patients with high or low fatigue Whole blood 251 differentially methylated CpG sites were associated with fatigue. The CpG site with the most pronounced hypomethylation in pSS high fatigue annotated to the SBF2-antisense RNA1 gene. The most distinct hypermethylation was observed at a CpG site annotated to the lymphotoxin alpha gene.
France
Thabet Y et al.
(66)
Methylation analysis and transcript levels of DNMTs in patients with and without SS Peripheral B and T cells; SGECs Global demethylation and reduction in DNMT1 transcript levels in SGECs of SS patients. No differences in methylation profile for B and T cells.
France
Gestermann N et al.
(105)
Methylation analysis of IRF5 promoter region to determine if this could be the cause for increased IRF5 mRNA expression in patients with SS Peripheral B and T cells DNA methylation profile of T CD4+ and B lymphocytes did not differ between SS patients and controls.
France
Miceli-Richard C et al.
(75)
Methylation comparison in profiles in the CD4+ T cells and CD19+ B cells of pSS patients and controls CD19+ B cells 55% of genes had hypermethylated CpG sites, 38% had hypomethylated CpG sites, and the remaining 7% of genes exhibited a mixed methylation pattern at CpG sites in CD19+ B cells. Genes with significantly different DNA methylation patterns are involved in functional pathways required for inflammation.
China
Yu X et al.
(68)
Methylation analysis of FOXP3 promoter region to determine whether the FOXP3 promoter is hypermethylated and whether aberrant FOXP3 expression occurs in CD4+ T cells from patients with pSS CD4+ T cells Hypermethylation at the promoter of FOXP3 in CD4+T cells of pSS patients. A decrease in expression in protein FOXP3 mRNA and protein in CD4+T cells of pSS patients.
China
Yin H et al.
(107)
Methylation analysis of TNFSF7 promoter region CD4+ T cells Hypermethylation at the promoter of TNFSF7 in pSS CD4+T cells. Demethylation of the CD70 promoter regulatory elements contributes to CD70 overexpression in pSS CD4+ T cells and may contribute to autoreactivity.
France
Belkhir R et al.
(108)
methylation analysis of CD40L
promoter region
CD4+ T cell No significant differences in methylation profiles between patients and controls.
Greece
Mavragani CP et al.
(109)
Methylation analysis of LINE-1; expression analysis of DNMT1, DNMT3A, DNMT3B, MeCP2 in SS patients and controls SGECs ↑ levels of DNMT1, DNMT3B and MeCP2 transcripts in SS patients.
France
Konsta OD et al.
(110)
Methylation analysis at SSB promoter region in patients with pSS and cell cultures SGECs ↓ global DNA methylation levels in patients with SS. ↓ global methylation associated with lymphocyte infiltration in MiSG; ↓ global methylation associated with anti-La/SSB positive SS cases demethylation at SSB promoter caused higher levels of transcripts and SSB super expression in 5-Aza-C-treated HSG cells.
France
Konsta OD et al.
(111)
Methylation analysis of the KRT19 locus to investigate epigenetic regulation of expression in patients with SS SGECs ↓ global DNA methylation in pSS patients is associated with demethylation of the KRT19 locus as well as with overexpression of the KRT19 protein.
USA
Chi C et al.
(112)
Methylation analysis in patients with and without SS SGECs 215 DMRs in SS patients: 169 hypermethylated regions related to nervous system development, cell signaling and transport; and 46 hypomethylated regions related to immune function.
USA
Cole MB et al.
(113)
Methylation analysis in patients with varying phenotypes of SS SGECs 7,820 DMPs associated with disease status (5,699 hypomethylated and 2,121 hypermethylated DMPs); 57 of the genes with DMPs are involved with the immune response; extensive hypomethylated region near genes PSMB8 and TAP1.
Chile
Sepúlveda D et al.
(114)
Methylation and expression analysis in genes of the IRE1α/XBP-1 pathway in SS and control patients SGECs Hypermethylation in IRE1α, XBP-1 and GRP78
promoter region and diminished transcript levels; ↓ protein levels for IRE1α, XBP-1s and GRP78 in SS patients.

SS, Sjögren’s syndrome; pSS, Primary Sjögren’s syndrome; SGECs, salivary gland epithelial cells; CpG sites, cytosine guanine dinucleotide (CpG) sites; DNMTs, DNA methyltransferase 1; MiSG, minor salivary gland; FOXP3, fork head Box Protein 3; LINE-1, long interspersed repeat element 1; anti, SSB, anti-Sjögren’s syndrome antigen B; HSG, human salivary gland; KRT19, keratin 19; DMRs, Differentially methylated regions; DMPs, differentially methylated positions; IRE1α, inositol-requiring enzyme 1alpha; XBP-1s, X-box binding protein 1s; GRP78, glucose-regulating protein 78.