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. Author manuscript; available in PMC: 2024 Nov 1.
Published in final edited form as: Nat Rev Cardiol. 2023 Nov 20;21(5):326–345. doi: 10.1038/s41569-023-00952-5

Table 2 |.

lncRNAs that suppress cardiac hypertrophy and ameliorate cardiac function

lncRNA Models Functions Interactions and effects Ref.
Ahit (antihypertrophic interrelated transcript) Mouse and in vitro (rat cardiomyocytes) Inhibition of Ahit induces cardiac hypertrophy, both in vitro and in vivo Interacts with SUZ12 163
Caren (cardiomyocyte-enriched non-coding transcript) Mouse Cardioprotective effects by regulating the translation of a distant gene and maintaining cardiomyocyte homeostasis Interacts with HINT1 49
CHAR (cardiac hypertrophy-associated regulator) Mouse and in vitro (cardiomyocytes) CHAR downregulation is sufficient to induce hypertrophic phenotypes in healthy mice; overexpression of CHAR reduced the hypertrophic response Interacts with miRNA-20b (downregulation) 164
CPhar (cardiac physiological hypertrophy-associated regulator) Mouse (exercise model) Overexpression of CPhar prevents myocardial ischaemia-reperfusion injury and cardiac dysfunction in vivo CPhar works with DDX17 as a binding partner to sequester C/EBPβ, leading to decreased levels of ATF7 165
GAS5 (growth arrest-specific 5) Mouse (Srsf4 knock-out) Represses cardiac hypertrophy in Srsf4 knock-out mice GAS5 is a repressor of the glucocorticoid receptor 166
H19 Human and mouse H19 inhibits E3 ligase-dependent polyubiquitination at Lys3584 of dystrophin (referred to as Ub-DMD), impeding dystrophin protein degradation Interacts with dystrophin 71,72
In vitro (cardiomyocytes) H19–miRNR-675 axis acts as a negative regulator of cardiac hypertrophy by targeting CaMKIIδ Inhibition of miRNA-675 reversed the increase in cardiomyocyte size associated with H19 overexpression; implicates miRNA-675 derived from H19 in cardiac hypertrophy 68,167
lnccytb (lncRNA cytb) Mouse Cytosolic lncRNA; function consistent with microRNA sponge Interacts with miRNA-103–3p 168
lncMYH7b In vitro (induced pluripotent stem cell-derived cardiomyocytes) lncMYH7b regulates the ratio of MHCβ to MHCα, thereby influencing cardiac rhythm Interacts with TEAD3 126
Mhrt or Myheart (myosin heavy-chain-associated RNA transcripts) Mouse Highly expressed in adult heart; stress represses Mhrt expression; overexpression of Mhrt protects the heart against cardiac hypertrophy and heart failure in response to stress Mhrt antagonizes the function of the chromatin remodelling enzyme BRG1 117
Mhrt (lncRNA-Mhrt) Mouse and in vitro (cardiomyocytes) lncRNA-Mhrt inhibits cardiac hypertrophy by inhibiting myocardin Interacts with myocardin through miRNA-145a-5p 169
Mhrt779 Mouse (exercise and TAC models) Mhrt779 increases antihypertrophic effects associated with other interventions Interacts with BRG1-HDAC2 120122
OIP5-AS1 (Opa interacting protein 5-antisense RNA 1; 1700020I14Rik; Cyrano) Mouse (knock-out model) Heart failure associated with TAC-induced pressure overload is exacerbated in female (but not male) knock-out mice Interacts with genes involved with mitochondrial function 170
Trdn-as (triadin-antisense) Mouse and in vitro Role in cardiac conduction through splicing of target genes Interacts with triadin 45
ZNF593-as (RP11–96L14.7; ENST00000448923.2) Mouse Localized in the cytoplasm of cardiomyocytes; levels are decreased in the failing hearts of patients with dilated cardiomyopathy; improves TAC-induced cardiac dysfunction ZNF593-as acts as a guide or scaffold for HNRNPC with RYR2 mRNA, thereby stabilizing RYR2 mRNA 171

lncRNA, long non-coding RNA; miRNA, microRNA; TAC, transverse aortic constriction.