Figure 4: Population structure in people with a Yamnaya cultural affiliation.

Individuals are projected in the same space as in Fig. 1. (a) showing that the Core Yamnaya cluster (black symbols) from diverse sites is differentiated from the Don Yamnaya (blue) who tend towards the UNHG. (b) Yamnaya individuals in the West (Ukraine, Hungary, Slovakia, and Southeastern Europe) include a tight cluster of individuals as well as others that tend towards the direction of European Neolithic and Chalcolithic groups from Romania and Hungary. Individuals from Russia are shown in grey circles in panel (b).