Table 1.

Model parameters and variables

Parameter/variable	Description	Default value or range
$a$	Reproduction rate of adult hosts	n/a
$a_0$	Baseline reproduction rate of adult hosts	5
$b_J,b_A$	Natural mortality rate of juvenile/adult hosts	1/5
$b$	Natural mortality rate of all hosts when $b_J=b_A$	1/5
$c_1$	Strength of tolerance-reproduction trade-off	0.25
$c_2$	Shape of tolerance-reproduction trade-off	3
$g$	Host maturation rate	1
$I_J,I_A$	Density of infected juveniles/adults	n/a
$N$	Host population density	n/a
$q$	Strength of host density-dependence	1
$S_J,S_A$	Density of susceptible juveniles/adults	n/a
$t$	Time, measured in arbitrary units	n/a
${\tau }_A$	Adult tolerance	0
${\tau }_J,{\tau }_L$	Juvenile/lifelong tolerance	$0\le {\tau }_J,{\tau }_L\le 1$
$\alpha$	Mortality virulence	$0\le \mathrm{\alpha }$
${\beta }_0$	Baseline transmission rate	10
$\gamma$	Recovery rate	0 or 1
$\lambda$	Force of infection	n/a
$\varphi$	Relative pathogen mutation rate	20