Catabolic pathway of vanillate for S. paucimobilis SYK-6 via the PCA 4,5-cleavage pathway. LigA and LigB, small and large subunits of PCA 4,5-dioxygenase (4,5-PCD) (27, 38); LigC, CHMS dehydrogenase (this study); LigI, PDC hydrolase (23); LigH, an essential gene product for vanillate and syringate O demethylations (26).