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Tetrasaccharide linker biosynthesis. UXS1 catalyzes the synthesis of UDP‐xylose from UDP‐glucuronic acid, and the glycosyltransferases XYLT1 and XYLT2 transfer the xylosyl group to a serine residue on the core protein. B4GALT7, B3GALT6 and B3GAT3 then add a β1,4‐linked galactose, a β1,3‐linked galactose and a β1,3‐linked glucuronic acid, respectively.
