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. 2024 May 29;43(14):3009–3026. doi: 10.1038/s44318-024-00128-y

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© The Author(s) 2024

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. Creative Commons Public Domain Dedication waiver http://creativecommons.org/publicdomain/zero/1.0/ applies to the data associated with this article, unless otherwise stated in a credit line to the data, but does not extend to the graphical or creative elements of illustrations, charts, or figures. This waiver removes legal barriers to the re-use and mining of research data. According to standard scholarly practice, it is recommended to provide appropriate citation and attribution whenever technically possible.

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(A) C4BP α7β1 (C4BP∆ProS sample). A central core with a diameter of ~5 nm surrounded by 7 C4BPα “arms” and a shorter C4BPβ chain. The C4BPα, C4BPβ, and 8 CCP domains of α2 were annotated. (B) Representative images of the four major species identified in human serum C4BP sample (α6β1 and α7β1 variants with or without ProS). (C) HS-AFM time series following a single C4BP molecule in a strong immobilization buffer highlighting the flexibility of C4BPα. Domains moving between frames are indicated by blue arrows. All scale bars correspond to 20 nm. Source data are available online for this figure.