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. 2024 Jul 15;15:1419645. doi: 10.3389/fmicb.2024.1419645

Table 2.

Classification of soil phosphorus cycling microbial functional genes.

Classification KO number Gene Reference
I. P-starvation response regulation
1. P-starvation response regulation
(1) Phosphate regulon sensor histidine kinase K07636 phoR Choi et al. (2009) and Liu J. et al. (2018)
(2) Phosphate regulon response regulator K07657 phoB Choi et al. (2009) and Liu J. et al. (2018)
(3) Negative regulator of PhoR/PhoB two-component regulator K02039 phoU Choi et al. (2009) and Liu J. et al. (2018)
II. P-uptake and transportation
2. Inorganic phosphate transportation
(4) Phosphate transport system ATP-binding protein K02036 pstB Liu J. et al. (2018)
(5) Phosphate transport system permease protein K02037 pstC Liu J. et al. (2018)
(6) Phosphate transport system permease protein K02038 pstA Liu J. et al. (2018)
(7) Phosphate transport system substrate-binding protein K02040 pstS Liu J. et al. (2018)
(8) Inorganic phosphate transporter K03306 pit Liu J. et al. (2018)
3. Phosphate transportation
(9) sn-glycerol 3-phosphate transport system substrate-binding protein K05813 ugpB Liang et al. (2020)
(10) sn-glycerol 3-phosphate transport system permease protein K05814 ugpA Liang et al. (2020)
(11) sn-glycerol 3-phosphate transport system permease protein K05815 ugpE Liang et al. (2020)
(12) sn-glycerol 3-phosphate transport system ATP-binding protein K05816 ugpC Liang et al. (2020)
(13) Glycerol uptake operon antiterminator K02433 glpP Beijer and Rutberg (1992)
(14) Glycerol-3-phosphate transporter K02445 glpT Beijer and Rutberg (1992)
4. Phosphonate transportation
(15) Phosphonate transport system ATP-binding protein K02041 phnC Liu J. et al. (2018)
(16) Phosphonate transport system permease protein K02042 phnE Liu J. et al. (2018)
(17) Phosphonate transport system substrate-binding protein K02044 phnD Liu J. et al. (2018)
(18) 2-amino-ethyl phosphonate transport system permease protein K11082 phnV Jiang et al. (1995)
III. Inorganic P-solubilization and organic P mineralization
5. Inorganic phosphate solubilization
(19) Quinoprotein glucose dehydrogenase gcd K00117 gcd Andreeva et al. (2011) and An and Moe (2016)
(20) Polyphosphate kinase K00937 ppk Liu J. et al. (2018)
(21) Inorganic pyrophosphatase K01507 ppa Liu J. et al. (2018)
(22) Exopolyphosphatase K01524 ppx Liu J. et al. (2018)
(23) B Pyrroloquinoline quinone biosynthesis protein B K06136 pqqB Andreeva et al. (2011) and An and Moe (2016)
(24) Pyrroloquinoline-quinone synthase K06137 pqqC Andreeva et al. (2011) and An and Moe (2016)
(25) Pyrroloquinoline quinone biosynthesis protein K06138 pqqD Andreeva et al. (2011) and An and Moe (2016)
(26) PqqA peptide cyclase K06139 pqqE Andreeva et al. (2011) and An and Moe (2016)
6. Phosphate ester mineralization
(27) Alpha-glycerophosphate oxidase K00105 glpO Beijer and Rutberg (1992)
(28) Glycerol-3-phosphate dehydrogenase K00111 glpA Beijer and Rutberg (1992)
(29) Glycerol-3-phosphate dehydrogenase subunit B K00112 glpB Beijer and Rutberg (1992)
(30) Glycerol-3-phosphate dehydrogenase subunit C K00113 glpC Beijer and Rutberg (1992)
(31) Glycerol kinase K00864 glpK Beijer and Rutberg (1992)
(32) Glycerol uptake facilitator protein K02440 glpF Beijer and Rutberg (1992)
(33) Glycerol-3-phosphate regulon repressor K02444 glpR Beijer and Rutberg (1992)
(34) Alkaline phosphatase K01077 phoA Liu J. et al. (2018)
(35) Alkaline phosphatase D K01113 phoD Liu J. et al. (2018)
(36) Acid phosphatase K01078 olpA Liu J. et al. (2018)
(37)(A)Acid phosphatase (class A) K09474 phoN Liu J. et al. (2018)
(38) 3-phytase K01083 phyA Liu J. et al. (2018)
(39) 4-phytase/acid phosphatase K01093 appA Liu J. et al. (2018)
(40) Phosphotriesterase-related protein K07048 opd Beijer and Rutberg (1992)
(41) Glycerophosphoryl diester phosphodiesterase K01126 ugpQ Beijer and Rutberg (1992)
7. Phosphonate mineralization
(42) C-P lyase core complex, Phosphonate transport system regulatory protein K02043 phnF Liu J. et al. (2018)
(43) C-P lyase core complex, Ribose 1,5-bisphosphokinase K05774 phnN Liu J. et al. (2018)
(44) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate synthase subunit K05780 phnL Liu J. et al. (2018)
(45) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate synthase subunit K05781 phnK Liu J. et al. (2018)
(46) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate diphosphatase K06162 phnM Liu J. et al. (2018)
(47) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate synthase subunit K06163 phnJ Liu J. et al. (2018)
(48) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate synthase subunit K06164 phnI Liu J. et al. (2018)
(49) C-P lyase core complex, Alpha-D-ribose 1-methylphosphonate 5-triphosphate synthase subunit K06165 phnH Liu J. et al. (2018)
(50) C-P lyase core complex, Alpha-D-ribose 1- methylphosphonate 5-triphosphate synthase subunit K06166 phnG Liu J. et al. (2018)
(51) C-P lyase core complex, Phosphoribosyl 1,2-cyclic phosphate phosphodiesterase K06167 phnP Liu J. et al. (2018)
(52) C-P lyase core complex, Aminoalkylphosphonate N-acetyltransferase K09994 phnO Liu J. et al. (2018)
(53) Alkylphosphonate utilization operon protein K06193 phnA Liu J. et al. (2018)
(54) 2-aminoethylphosphonate-pyruvate transaminase K03430 phnW Liu J. et al. (2018)
(55) Phosphonoacetaldehyde hydrolase K05306 phnX Liu J. et al. (2018)