A simplified model of evolutionary events in Brassicaceae.
After splitting from Aethionemeae, the progenitors of core Brassicaceae (colored triangles) diversified in a short period between 19.3 and 24.4 mya during the late Oligocene to the early Miocene. The first random duplication of TPS4s occurred after the At-α WGD. Two independent random duplication events doubled the TPS2-likes and TPS2s in Hesperodae and the MRCA of other supertribes (Brassicodae, Heliophilodae, Camelinodae, and Arabodae), respectively. The deep-sea temperatures are modified from Zachos et al. (2001).