IMD2 mRNA synthesis is regulated by GTP-dependent start site selection and transcriptional attenuation. Imd2 catalyzes the first committed step of GTP synthesis from IMP, which is also a precursor of ATP. IMD2 has multiple transcription start sites (TSS, bent arrows). Upstream TSS are used when cellular GTP is sufficient, producing transcripts containing a Nrd1-Nab3-Sen1 (NNS) terminator that terminate prematurely and are degraded by the nuclear exosome. When cellular GTP is deficient, a downstream TSS is preferentially used, resulting in full-length mRNA [adapted from reference (8)].