These studies were carried out in the following assay buffers at room temperature: sodium acetate (pH 5.2 and 5.6), Hepes (pH 6.0, 6.5, 7.1 and 7.4) and Tris/HCl (pH 8.0). Reactions were initiated by the addition of enzyme (50 pM final enzyme concentration) to solutions of buffer containing substrate (2–10 nM). The points are experimental and the lines are theoretical for the single ionization equation k=k~/(1+[H+]/Ka), where k=kcat or k=kcat/Km and k~=k~cat or k~cat/K~m (pH-independent kinetic parameters) (a) pKa=7.1±0.1, k~cat/K~m=(4.1±0.1)× 105 M−1·s−1 and (b) pKa=6.6±0.1, k~cat=0.009±0.001 s−1.