Fig. 22. CanDo example of DNA skips and strain compensation on a square lattice for a planar origami. The top row shows the handedness of the torque as determined by the presence and location of skips. Red indicates right-handed torque helices, and blue indicates left-handed torque in the twist schematic. The average twist values are indicated for each of the four designs. As noted in the text, the number of base pairs per turn in an unstrained dsDNA helix is 10.5. The bottom row shows the outputs of the corresponding CanDo models. These illustrate the approximate shape and local helix mobility.

Twist Schematic
Average Twist Values	Default Length: slightly overwound 10.66 bp/turn 33.75A/bp	Alternating Skips: 10.333 bp/turn 34.84A/bp	A1/4 Skips: 10.5 bp/turn 34.28A/bp	A1/4 Skips staggered: 10.5 bp/turn 34.28A/bp
CanDo Prediction