Metastatic
state of MDA-MB-231 cells influences cellular viscoelasticity
revealed by QCM-D. (a) Illustration of QCM-D cell-based experiments
in liquid conditions and the corresponding plots reporting changes
in frequency (ΔF) and dissipation energy (ΔD) vs time according to the rigidity or
softness of the cell layer. QCM-D utilizes a converse piezoelectric
effect: the AT-cut quartz crystal oscillates when an electric voltage
is applied. The mechanical oscillations can be transferred into an
equivalent electric circuit model, which allows the depiction of a
complete description of the oscillation in the presence of a mass
and viscous loading. In the case of soft and viscoelastic cell monolayers,
the amplitude of the crystal oscillation decreases rapidly due to
the high energy losses compared to rigid cell monolayers. (b–j)
Real-time QCM-D measurements. (b) ΔF- and (c)
ΔD-responses, and the corresponding (d) Df -plots with linear regression of low cell count MDA-MB-231
cells with varying expression of Cx43 undergoing cell detachment.
(e) ΔF- and (f) ΔD-responses
and the corresponding (g) Df-plots with linear regression
of high cell count MDA-MB-231 cells with varying expression of Cx43
undergoing cell detachment. The cells were seeded on top of AT-cut
gold-coated quartz sensors for 24 h, after which they were mounted
on the modules and washed with PBS for 10 min at a 10 μL/min
flow rate. The cells were detached with trypsin at 30 μL/min,
and the frequency and dissipation were recorded over a period of 35
min. The cells were then washed with PBS to remove the unbound cells.
Change in (h) frequency and (i) dissipation amplitudes and (j) Df-slopes for MDA-MB-231 cells with varying Cx43 expression
at different cell count states. The values depicted are the mean ±
SEM from at least three separate experiments evaluated per condition.
*** denotes a p-value <0.001, ** denotes a p-value <0.01, and * denotes a p-value
<0.05 compared to different conditions using ANOVA followed by
the post hoc Tukey HSD test. The data shown are from the same control
and experimental pair.