Fossil site paleo‐precipitation reconstructions for seven paleocommunities including, (1) January Cave, (2) Little Box Elder Cave, (3) Brynjulfson Cave 1, (4) Anderson Pit, (5) New Trout Cave, (6) McKittrick, and (7) Friesenhahn Cave. (a) Observed annual precipitation millimeters natural log‐transformation (AP). Fossil sites are numbered and indicated as black circles. (b) Trait turnover of RBL at each fossil site represented in ecometric space. (c) Observed mean annual temperature (MAT). Fossil sites are numbered and indicated as black circles. (d) Trait turnover of RBL at each fossil site represented in ecometric space. Ecometric space is based on the RBL and maximum likelihood estimate of AP, where each grid cell represents communities that display a given RBL mean and standard deviation and are color‐coded based on the maximum likelihood estimate of AP or MAT. Paleocommunities are represented in the ecometric space as numbered circles and arrows indicate the ecometric space inhabited by the modern communities located in the same geographic location.