TABLE 4.

Nucleotide diversity of geographically different CTV populations^a

Group	Region	Nucleotide diversity			Vb				Vw	Nst
		Sp		Cal	Sp-Cal
1	A	0.17627 (0.08425)		0.05809 (0.01306)	0.00819 (0.01924)				0.11718 (0.05909)	0.06496
	F	0.06825 (0.02866)		0.03328 (0.01094)	0.00182 (0.00764)				0.04846 (0.01979)	0.03620
	C	0.03518 (0.00749)		0.03264 (0.01025)	0.00059 (0.00380)				0.03391 (0.00582)	0.01710
	P	0.05390 (0.01083)		0.05349 (0.01174)	0.00351 (0.00537)				0.05369 (0.00784)	0.06136
2		Sp	Cal	Por	Sp-Cal	Cal-Por	Sp-Por	Sp-Cal-Por	Vw	Nst
	C	0.03518	0.03264	0.06946	0.00059	0.00432	0.00209	0.00233	0.04576	0.04845
		(0.00749)	(0.01025)	(0.01535)	(0.00380)	(0.01103)	(0.00965)	(0.00722)	(0.01187)
3		Ja		Por	Ja-Por				VW	Nst
	C2	0.08262 (0.00652)		0.07798 (0.01236)	0.00544 (0.00798)				0.0803 (0.00608)	0.06344

^aGenomic regions A, F, C (nt 424 to 647 of the coat protein gene), P (Table 1 and Fig. 1), and C2 (first 462 nt of coat protein gene) were analyzed. The first group shows diversities of regions A, F, C, and P using the sequences determined by us here. The second includes comparisons only of region C, using our sequences and additional nucleotide sequences obtained from GenBank (see Materials and Methods). The last shows nucleotide diversities of region C2 estimated from sequences obtained from GenBank. The CTV isolates analyzed were from Spain (Sp), California (Cal), Portugal (Por), and Japan (Ja). V_b, average nucleotide diversity between geographic populations (24); V_w, average nucleotide diversity within geographic populations (24); N_st = V_b/(V_w + V_b), proportion of the diversity between geographic populations with respect to total nucleotide diversity (24). Nucleotide diversity is the average number of nucleotide substitutions per site in each pair of sequence variants. Standard errors are indicated in parentheses.