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Example records showing measurement of bidirectional sliding friction. (A–C) Force and position are plotted against time for beads decorated with (A) native yeast kinetochore particles, (B) recombinant yeast Ndc80c, or (C) recombinant human Ndc80c, attached to the sides of coverslip-anchored microtubules and pulled alternately toward the plus (red traces) and minus ends (blue traces). Mean forces and speeds for each sliding event are indicated.
