Identification of codon sites evolving under positive selection in natural populations of VSIV. The figure shows the 42 codons under positive selection identified at multiple genes VSIV by MEME analysis. α: synonymous substitution rate, β1:Non-synonymous substitution rate for the negative/neutral evolution component 1, p1: mixture distribution weight allocated to negative/neutral evolution component 1, β+:non-synonymous substitution rate at a site for the positive selection component, p+:mixture distribution weight allocated to the positive selection component, LTR: likelihood test statistics for episodic diversification, i.e., p+ > 0, p-value: asymptotic p-value for episodic diversification, i.e., p+ > 0, # branches: the (very approximate and rough) estimate of how many branches have been under selection at this site, i.e., had an empirical Bayes factor of 100 or more for the β+ rate, q: and class: selection kind.