1 |
Male mandibles with dorsal tooth |
2
|
– |
Male mandibles without dorsal tooth |
9
|
2 |
Internal sac of aedeagus with two spines |
3
|
– |
Internal sac of aedeagus with one spine |
4
|
3 |
HL/PL more than 1.00. Male right mandible with small dorsal tooth; internal sac of aedeagus with one long curved spine and one short spine. Posterior margin of female tergite VIII truncate |
P. (Gnathopaederus) szechuanus (Chapin, 1927)
|
– |
HL/PL no more than 0.95. Male right mandible with large dorsal tooth; internal sac of aedeagus with two short spines. Posterior margin of female tergite VIII convex |
P. (Gnathopaederus) bursavacua Willers, 2001
|
4 |
Smaller species, length of body: 7.2 mm. Length of antenna no more than 3.0 mm |
P. (Gnathopaederus) jilongensis Li & Zhou, 2009
|
– |
Larger species, length of body more than 8.0 mm. Length of antenna more than 3.3 mm |
5
|
5 |
Male sternite V–VI with shallow median impression posteriorly, this impression with weakly modified setae |
P. (Gnathopaederus) zhangmuensis Cheng & Peng, 2019
|
– |
Male sternite V–VI without median impression and modified setae posteriorly |
6
|
6 |
Dorsal plate of aedeagus straight in lateral view and internal sac with one short spine |
P. (Gnathopaederus) cheni Peng & Li, 2015
|
– |
Dorsal plate of aedeagus weakly curved in lateral view and internal sac with one long spine |
7
|
7 |
Male left mandible with large dorsal tooth; internal spine of aedeagus apically extending nearly to apex of dorsal plate |
P. (Gnathopaederus) furcillatus Assing, 2017
|
– |
Male left mandible with small dorsal tooth; internal spine of aedeagus far from reaching apex of dorsal plate |
8
|
8 |
Smaller species, length of body: 8.3–10.5 mm. Male right mandible with small dorsal tooth; smaller aedeagus with slenderer parameres |
P. (Gnathopaederus) yunnanensis Willers, 2001
|
– |
Larger species, length of body 11.6–12.4 mm. Male right mandible with large dorsal tooth; larger aedeagus with stouter parameres |
P. (Gnathopaederus) xuei Peng & Li, 2015
|
9 |
Coloration of abdomen black |
10
|
– |
Abdomen bicoloured (usually segments III–VI reddish and segments VII–X black) |
21
|
10 |
Coloration of head brown. Male sternite VI with distinct median impression posteriorly |
P.lateralis Li, Solodovnikov & Zhou, 2014
|
– |
Coloration of head black. Male sternite VI without median impression posteriorly |
11
|
11 |
Male sternite VII with deep median impression posteriorly. Parameres of aedeagus very short |
P.mirus Yang & Peng, sp. nov.
|
– |
Male sternite VII without impression posteriorly. Parameres long and conspicuous |
12
|
12 |
Small species, length of body no more than 8.5 mm |
P.sinisterobliquus Li, Zhou & Solodovnikov, 2013
|
– |
Large species, length of body larger than 9.0 mm |
13
|
13 |
Maximum width of pronotum no more than 1.50 mm. Parameres of aedeagus apically distinctly curved in lateral view |
P.chentangus Yang & Peng, sp. nov.
|
– |
Maximum width of pronotum more than 1.54 mm. Parameres of aedeagus apically weakly curved or straight in lateral view |
14
|
14 |
Internal sac of aedeagus with two spines |
15
|
– |
Internal sac of aedeagus with three spines |
17
|
15 |
Maximum width of abdomen more than 2.1 mm. Parameres of aedeagus slender. Female sternite IX longer |
P.parvidenticulatus Li, Zhou & Solodovnikov, 2013
|
– |
Maximum width of abdomen no more than 2.0 mm. Parameres of aedeagus stout. Female sternite IX shorter |
16
|
16 |
HL/HW more than 0.95. Aedeagus with apically hooked dorsal plate in dorsal view and two long sclerotized spines in internal sac. Female sternite VIII with elliptic depression posteriorly |
P.volutobliquus Li, Zhou & Solodovnikov, 2013
|
– |
HL/HW no more than 0.90. Aedeagus with apically acute dorsal plate in dorsal view and two short sclerotized spines in internal sac. Female sternite VIII without depression |
P.nanlingensis Peng & Li, 2016
|
17 |
Legs with infuscate apical portion of femora. Aedeagus with apically hooked dorsal plate in dorsal view |
P.biacutus Li, Zhou & Solodovnikov, 2013
|
– |
Legs with brown femora. Aedeagus with apically acute dorsal plate in dorsal view |
18
|
18 |
Parameres of aedeagus symmetric and slender in ventral view |
P.jianyueae Peng & Li, 2014
|
– |
Parameres of aedeagus asymmetric and stouter in ventral view |
19
|
19 |
Internal sac of aedeagus with one additional dark membranous structure. Female right mandible with single-pointed middle tooth |
P.zhaoi Yang & Peng, sp. nov.
|
– |
Internal sac of aedeagus without dark membranous structure. Female right mandible with small bifid molar tooth apically |
20
|
20 |
Ventral plate of aedeagus short and broad. Posterior margin of female sternite VIII strongly convex |
P.songi Yang & Peng, sp. nov.
|
– |
Ventral plate of aedeagus long and slender. Posterior margin of female sternite VIII distinctly trifurcate |
P.trispinosus Yang & Peng, sp. nov.
|
21 |
Aedeagus stout, with short and stout dorsal plate in dorsal view |
22
|
– |
Aedeagus slender, with long and slender dorsal plate in dorsal view |
24
|
22 |
Legs reddish with infuscate apical portion of femora. Aedeagus with apically acute dorsal plate in dorsal view |
P.tibetanus Cameron, 1928
|
– |
Legs yellowish brown. Aedeagus with apically convex dorsal plate in dorsal view |
23
|
23 |
Male labrum with sinuate anterior margin. Aedeagus with short parameres. Female sternite VIII with truncate median process posteriorly |
P.describendus Willers, 2001
|
– |
Male labrum with deeply excavate anterior margin. Aedeagus with long parameres. Posterior margin of female sternite VIII convex |
P.daicongchaoi Peng & Li, 2016
|
24 |
Dorsal plate of aedeagus not reaching apices of parameres |
25
|
– |
Dorsal plate of aedeagus extending beyond apices of parameres |
29
|
25 |
Segments III–VI of abdomen reddish and with black patch in middle; elytra with pronounced impression |
P. (Harpopaederus) gottschei Kolbe, 1886
|
– |
Segments III-VI of abdomen pale-reddish and without black patch in middle; elytra without impression |
26
|
26 |
Dorsal plate of aedeagus without denticles |
P. (Harpopaederus) antennocinctus Willers, 2001
|
– |
Dorsal plate of aedeagus with conspicuous denticles |
27
|
27 |
Internal sac of aedeagus with one long and apically acute moderately sclerotized structure |
P. (Harpopaederus) willersi Assing, 2020
|
– |
Internal sac of aedeagus with one basal clip-shaped structure and one asymmetric apical structure |
28
|
28 |
Length of aedeagus: 2.4 mm. Female tergite VIII apically narrower |
P. (Harpopaederus) deplectens Assing, 2015
|
– |
Length of aedeagus: 2.7 mm. Female tergite VIII apically broader |
P. (Harpopaederus) chinensis Bernhauer, 1931
|
29 |
Dorsal plate of aedeagus without denticles |
30
|
– |
Dorsal plate of aedeagus with denticles |
31
|
30 |
Length of aedeagus: 1.4 mm; internal sac without sclerotized basal structures |
P. (Harpopaederus) xui Peng & Li, 2015
|
– |
Length of aedeagus: 2.0 mm; internal sac with one clip-shaped, weakly sclerotized basal structure |
P. (Harpopaederus) edentulus Assing, 2015
|
31 |
Legs with blackish metatibiae |
32
|
– |
Coloration of metatibiae much paler (usually yellowish) |
33
|
32 |
Length of aedeagus: 2.4–2.7 mm, with apically stouter dorsal plate. Female sternite VIII with short median process posteriorly |
P. (Harpopaederus) apfelsinicus Willers, 2001
|
– |
Length of aedeagus: 2.9 mm, with apically more slender dorsal plate. Female sternite VIII with long median process posteriorly |
P. (Harpopaederus) lineodenticulatus Li & Zhou, 2007
|
33 |
Forebody longer than 5.8 mm. Aedeagus conspicuously long (2.7 mm) |
P. (Harpopaederus) minicus Assing, 2015
|
– |
Forebody no more than 5.5 mm. Aedeagus shorter |
34
|
34 |
Internal sac of aedeagus with one long, apically acute and sclerotized apical structure |
35
|
– |
Internal sac of aedeagus without distinctly sclerotized apical structure |
38
|
35 |
Aedeagus 2.4 mm long, with longer apical portion of the dorsal plate |
P. (Harpopaederus) cultellatus Assing, 2015
|
– |
Aedeagus 1.9–2.1 mm long, with shorter apical portion of the dorsal plate |
36
|
36 |
Dorsal plate of aedeagus with hooked apex. Posterior margin of female sternite VIII trifurcate |
P. (Harpopaederus) yei Yang & Peng, sp. nov.
|
– |
Dorsal plate of aedeagus with acute apex. Female sternite VIII posteriorly with median process of triangular shape |
37
|
37 |
Tibiae usually yellowish; mandibles and shape of head without sexual dimorphisms. Aedeagus with moderately sclerotized apical internal structure |
P. (Harpopaederus) agnatus Eppelsheim, 1889
|
– |
Tibiae distinctly infuscate basally; mandibles and shape of head with pronounced sexual dimorphisms. Aedeagus with strongly sclerotized apical internal structure |
P. (Harpopaederus) konfuzius Willers, 2001
|
38 |
Femora brown. Dorsal plate of aedeagus with 20 small denticles; parameres apically weakly curved in lateral view |
P. (Harpopaederus) multidenticulatus Li, Zhou & Solodovnikov, 2014
|
– |
Femora bicoloured. Dorsal plate of aedeagus with several large denticles; parameres apically hooked in lateral view |
39
|
39 |
Head transverse (HL/HW: 0.90). Dorsal plate of aedeagus with short apical portion. Female sternite VIII with strongly convex posterior margin |
P. (Harpopaederus) brevior Li, Zhou & Solodovnikov, 2014
|
– |
Head weakly transverse (HL/HW: 0.99). Dorsal plate of aedeagus with long apical portion. Female sternite VIII with long median process posteriorly |
P. (Harpopaederus) gracilacutus Li & Zhou, 2007
|